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Published: 2011-08-11
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On the need to follow rigorously the Rules of the Code for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus Trimeresurus Lacépède, 1804 (Reptilia: Squamata: Viperidae)

Reptiles & Amphibiens, UMR 7205 OSEB, Département Systématique et Evolution, CP 30, Muséum National d’Histoire Naturelle, 57 rue Cuvier, 75231 Paris Cedex 05, France
Society for Southeast Asian Herpetology, Im Sand 3, 69115 Heidelberg, Germany
Département Systématique et Evolution, CP 30, Muséum National d’Histoire Naturelle, 57 rue Cuvier, 75231 Paris Cedex 05, France
Reptilia Nomenclature Onomatophore Nucleospecies Nominal species Spelling Type species Prenucleospecies Trimeresurus Trimeresurus viridis Cryptelytrops Trimeresurus insularis New synonym

Abstract

This paper analyzes the consequences of the non-respect of the Rules of the Code to ascertain the valid subsequent designation of the nucleospecies (type species) of the nominal genus Trimeresurus Lacépède, 1804. The long accepted designation was invalid because it was based on a nominal species which was not among the prenucleospecies (originally included species) of the nominal genus. In contrast with the commonly accepted viewpoint which makes the Indian taxon Coluber gramineus Shaw, 1802 the nucleospecies of the genus, we show that this role is played by Trimeresurus viridis Lacépède, 1804, a species inhabiting the Lesser Sunda Islands and Timor and, as a nomen oblitum, a senior synonym of Trimeresurus albolabris insularis Kramer, 1977, a taxon now considered a distinct species. The important nomenclatural implications of this finding are discussed here, especially with regard to the recent splitting of the genus Trimeresurus. The generic nomen Trimeresurus should be associated with the Trimeresurus albolabris group of species currently placed in the genus or subgenus Cryptelytrops Cope, 1860. A lectophoront (lectotype) is selected and described for Trimeresurus viridis Lacépède, 1804. Coluber viridis Bechstein, 1802 is an invalid objective junior synonym of Coluber gramineus Shaw, 1802. The current content of the genus Trimeresurus and of its eight subgenera is provided. Some clarifications or improvements to the Code are suggested.

References

  1. Anonymous (International Commission on Zoological Nomenclature) (1987) Opinion 1463. De Lacépède, 1788-1789, Histoire Naturelle des Serpens and later editions: rejected as a non-binominal work. Bulletin of Zoological Nomenclature, 44 (4), 265–267.

    Anonymous (International Commission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. Fourth Edition adopted by the International Union of Biological Sciences. International Trust of Zoological Nomenclature, London, xxix + 306 pp.

    Amaral, A. (1926). Notas de ophiologia. 4.a Nota de nomenclatura ophiologica. Sobre a differenciação dos nomes genericos Lachesis, Trimeresurus e Bothrops. Revista do Museum Paulista, 14, 34–40.

    Anderson, J. (1879) Anatomical and zoological researches: comprising an account of the zoological results of the two expeditions to western Yunnan in 1868 and 1875; and a monograph of the two cetacean genera, Platanista and Orcella. First Volume–Text. B. Quaritch, London, xxv + 985 pp.

    Barbour, T. (1912) A contribution to the zoögeography of the East Indian islands. Memoirs of the Museum of comparative Zoology, at Harvard College, 44 (1), 1–203.

    Bechstein, J.M. (1802) Herrn De la Cepede’s Naturgeschichte der Amphibien oder der eyerlegenden vierfüssigen Thiere und der Schlangen. Eine Fortsetzung von Büffon’s Naturgeschichte. Aus dem Französischen übersetzt und mit Anmerkungen und Zusätzen versehen von Johann Matthäus Bechstein. Vierter Band. Verlage des Industrie-Comptoir’s, Weimar, xx + 298 pp., Pl. 1–48.

    Beddome, R.H. (1862) Notes upon the land and fresh water snakes of the Madras Presidency. Madras quarterly Journal of medical Science, 5 (1), 1–32, Pl. 2.

    Bethencourt Ferreira, J. (1898) Reptis de Timôr no Museu de Lisboa. Jornal de Sciencias mathematicas, physicas e naturaes, (2), 5 (19), 151–156.

    Bleeker, P. (1860) Over de Reptiliën-fauna van Sumatra. Natuurkundig Tijdschrift voor Nederlandsch Indië, (5, Part I), 21, 284–298.

    Bleeker, P. (1861) Reptiliën van het eiland Timor. Natuurkundig Tijdschrift voor Nederlandsch Indië, (5, Part III), 22, 86–88.

    Blyth, E. (1846) Notes on the fauna of the Nicobar Islands. Journal of the Asiatic Society of Bengal, 15, 367–379.

    Blyth, E. (1861) Report of Curator, Zoological Department. Journal of the Asiatic Society of Bengal, 29 [1860] (1), 87–115.

    Boie, F. (1826) Generalübersicht der Familien und Gattungen der Ophidier. Isis von Oken, 19 (10), col. 981–982.

    Boie, F. (1827) Bemerkungen über Merrem’s Versuch eines Systems der Amphibien. Marburg. 1820. Erste Lieferung: Ophidier. Isis von Oken, 20 (10), col. 508–566.

    Note. Ulber (1993, Appendix: 323–380) provided a transcription in modern German alphabet and an English translation of this rare but fundamental publication.

    Bonaparte, C.-L. (1831) Saggio di una distribuzione metodica degli animali vertebrate. Presso Antonio Boulzaler, Rome, 1–78.

    Bonaparte, C.-L. (1832). Saggio d’una distribuzione metodica degli animali vertebrati a sangue freddo. Presso Antonio Boulzaler, Rome, 1–86.

    Bory de Saint-Vincent, J.-B. (1842) Traité élémentaire d’erpétologie ou Histoire naturelle des Reptiles, contenant des notions générales et particulières sur l’Antiquité, l’organisation, les mœurs de ces animaux, l’art de les conserver, avec la description et l’histoire de leurs familles et de leurs genres, d’après une méthode analytique; précédé d’une Introduction historique, et suivi d’une Biographie, d’une Bibliographie et d’un Vocabulaire. Mairet et Fournier, Paris, viii + 292 pp.

    Boulenger, G.A. (1890) The fauna of British India, including Ceylon and Burma. Reptilia and Batrachia. Taylor & Francis, London, xviii + 541 pp.

    Boulenger, G.A. (1896) Catalogue of the Snakes in the British Museum (Natural History). Volume III. Containing the Colubridae (Opisthoglyphae and Proteroglyphae), Amblycephalidae and Viperidae. British Museum (Natural History), London, xiv + 727 pp., Pl. 1–25.

    Boulenger, G.A. (1912) A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore, including the adjacent islands. Reptilia and Batrachia. Taylor & Francis, London, xiii + 294 pp.

    Bour, R. (2010) Constant Duméril’s Zoologie Analytique was published in 1805. Bionomina, 1, 56–57.

    Bour, R. (2011) François Marie Daudin (29 août 1776—30 novembre 1803), auteur de l’Histoire naturelle, générale et particulière, des Reptiles. Alytes, in press.

    Brattstrom, B.H. (1964) Evolution of the pit vipers. Transactions of the San Diego Society of natural History, 13 (11), 185–268.

    Brosse, J. (1998) Les tours du monde des explorateurs. Les grands voyages maritimes 1764–1843. Larousse, Paris, 288 pp.

    Burger, W.L. (1971) Genera of Pitvipers (Serpentes: Crotalidae). Ph.D. dissertation, University of Kansas, Lawrence (Kansas), 186 pp.

    Cantor, T.E. (1839) Spicilegium serpentium indicorum. Proceedings of the zoological Society of London, 1839, 31–34 + 49–55.

    Cantor, T.E. (1847) Catalogue of reptiles inhabiting the Malay Peninsula and islands, collected or observed by Theodore Cantor, Esq., M.D., Bengal Medical Service. Journal of the Asiatic Society of Bengal, 16, 1026–1078.

    Castoe, T.A. & Parkinson, C.L. (2006) Bayesian mixed models and the phylogeny of pitvipers (Viperidae: Serpentes). Molecular Phylogenetics and Evolution, 39 (1), 91–110.

    Catesby, M. (1743) The Natural History of Carolina, Florida and the Bahama Islands: Containing the Figures of Birds, Beasts, Fishes, Serpents, Insects, and Plants: particularly, the Forest-Trees, Shrubs, and other Plants, not hitherto described, or very incorrectly figured by Authors. Together with their Descriptions in English and French. To which are added Observations on the Air, Soil, and Waters: with Remarks upon Agriculture, Grain, Pulse, Roots, &c. To the whole is Prefixed a new and correct Map of the Countries treated of. Histoire naturelle de la Caroline, la Floride, & les Bahama: Contenant les Desseins des Oiseaux, Animaux, Poissons, Serpents, Insectes, & Plantes. Et en particulier, des Arbres des forets, Arbrisseaux, & autres plantes, qui n'ont point été decrits jusques à present par les Auteurs, ou peu exactement dessinés. Avec leurs Descriptions en François & en Anglois. A quoi on a adjouté des Observations sur l'Air, le Sol, & les Eaux, avec des Remarques sur l'Agriculture, les Grains, les Legumes, les Racines, &c. Le tout est precedé d'une Carte nouvelle & exacte des Païs dont il s'agist. Printed at the expense of the Author, London, (2), 100 pp., Pl. 1–100, xliv pp., (6).

    Cogger, H.G., Cameron, E.E. & Cogger, H.M. (1983) Zoological catalogue of Australia. Vol. 1. Amphibia and Reptilia. Australian Government Publishing Service, Canberra, vi + 313 pp.

    Cope, E.D. (1860) Catalogue of the venomous serpents in the museum of the Academy of Natural Sciences of Philadelphia, with notes on the families, genera, and species. Proceedings of the Academy of natural Sciences of Philadelphia, 11 [1859], 332–347.

    Creer, S., Malhotra, A. & Thorpe, R.S. (2003) Assessing the phylogenetic utility of four mitochondrial genes and a nuclear intron in the Asian pit viper genus, Trimeresurus: separate, simultaneous, and conditional data combination analyses. Molecular Biology and Evolution, 20 (8), 1240–1251.

    Creer, S., Pook, C.E., Malhotra, A. & Thorpe R.S. (2006) Optimal Intron Analyses in the Trimeresurus Radiation of Asian Pitvipers. Systematic Biology, 55 (1), 57–72.

    Cuvier, G. (1816) Le règne animal distribué d’après son organisation, pour servir de base à l’histoire naturelle des animaux et d’introduction à l’anatomie comparée. Tome II contenant les reptiles, les poissons, les mollusques et les annélidés. Déterville, Paris, xviii + 532 pp.

    Cuvier, G. (1829) Le règne animal distribué d’après son organisation, pour servir de base à l’histoire naturelle des animaux et d’introduction à l’anatomie comparée. Nouvelle édition, revue et augmentée. Tome II. Déterville, Paris, xv + 406 pp.

    Daudin, F.-M. (1803a) Division des ophidiens en vingt-trois genres. Bulletin des Sciences par la Société philomatique, 3 (72), 187–188.

    Daudin, F.-M. (1803b) Histoire naturelle, générale et particulière des reptiles. Ouvrage faisant suite aux oeuvres de Leclerc de Buffon, et partie du cours complet d’histoire naturelle rédigé par C. S. Sonnini, membre de plusieurs sociétés savantes. Tome sixième. Imprimerie F. Dufart, Paris, 447 pp., Pl. 71–80.

    David, P. & Ineich, I. (1999) Les serpents venimeux du monde: systématique et répartition. Dumerilia, Paris, 3, 3–499.

    David, P. & Vogel, G. (1996) The Snakes of Sumatra. An annotated checklist and key with natural history notes. Edition Chimaira, Frankfurt-am-Main, 260 pp.

    David, P., Vogel, G. & Vidal, N. (2003) On Trimeresurus fasciatus (Boulenger, 1896) (Serpentes: Crotalidae), with a discussion on its relationships based on morphological and molecular data. Raffles Bulletin of Zoology, 51 (1), 149–157.

    David, P., Vogel, G., Vijayakumar, S. P. & Vidal, N. (2006) A revision of the Trimeresurus puniceus-complex (Serpentes: Viperidae: Crotalinae) based on morphological and molecular data. Zootaxa, 1293, 1–80.

    Dawson, K., Malhotra, A., Thorpe, R.S., Guo, P., Mrinalini & Ziegler, T. (2008) Mitochondrial DNA analysis reveals a new member of the Asian pitviper genus Viridovipera (Serpentes: Viperidae: Crotalinae). Molecular Phylogenetics and Evolution, 49, 356–361.

    De la Cepède, B.G.E.L. (Comte de) (1789) Histoire naturelle des quadrupèdes ovipares et des serpens. Tome second. Imprimerie du Roi, Hôtel de Thou, Paris, 19 + 144 + 527 pp., Pl. 1–22.

    De Rooij, N. (1917) The reptiles of the Indo-Australian Archipelago. II. Ophidia. E. J. Brill, Leiden, xvi + 334 pp.

    Deraniyagala, P.E.P. (1955) A colored atlas of some vertebrates from Ceylon. Volume Three. Serpentoid Reptilia. Ceylon National Museums Publication, Colombo, xix + 121 pp., Pl. 1–48.

    Dowling, H.G. (1951) A proposed standard system of counting ventrals in snakes. British Journal of Herpetology, 1 (5), 97–99.

    Dubois, A. (1982) Le statut nomenclatural des noms génériques d’Amphibiens créés par Kuhl & Van Hasselt (1822): Megophrys, Occidozyga et Rhacophorus. Bulletin du Muséum national d’Histoire naturelle, (4), 4 (A), 261–280.

    Dubois, A. (1997) Proposals concerning the conditions needed for a name being eligible for conservation. Bulletin du Muséum national d’Histoire naturelle, (4), 18 (3–4), 317–320.

    Dubois, A. (2000) Synonymies and related lists in zoology: general proposals, with examples in herpetology. Dumerilia, 4 (2), 33–98.

    Dubois, A. (2005) Proposed Rules for the incorporation of nomina of higher-ranked zoological taxa in the International Code of Zoological Nomenclature. 1. Some general questions, concepts and terms of biological nomenclature. Zoosystema, 27 (2), 365–426.

    Dubois, A. (2008a) Authors of zoological publications and nomina are signatures, not persons. Zootaxa, 1771, 63–68.

    Dubois, A. (2008b) Phylogenetic hypotheses, taxa and nomina in zoology. In: A. Minelli, L. Bonato & G. Fusco (ed.), Updating the Linnaean heritage: names as tools for thinking about animals and plants, Zootaxa, 1950, 51–86.

    Dubois, A. (2010a) Retroactive changes should be introduced in the Code only with great care: problems related to the spellings of nomina. Zootaxa, 2426, 1–42.

    Dubois, A. (2010b) Zoological nomenclature in the century of extinctions: priority vs. “usage”. Organisms, Diversity & Evolution, 10, 259–274.

    Dubois, A. (2011) The International Code of Zoological Nomenclature must be drastically improved before it is too late. Bionomina, 2, 1–104.

    Dubois, A. & Bour, R. (2010) The nomenclatural status of the nomina of amphibians and reptiles created by Garsault (1764), with a parsimonious solution to an old nomenclatural problem regarding the genus Bufo (Amphibia, Anura), comments on the taxonomy of this genus, and comments on some nomina created by Laurenti (1768). Zootaxa, 2447: 1–52.

    Dubois, A. & Ohler, A. (1997) Early scientific names of Amphibia Anura. I. Introduction. Bulletin du Muséum national d’Histoire naturelle, (4), 18 (3–4), 297–320.

    Dubois, A. & Raffaëlli, J. (2009) A new ergotaxonomy of the family Salamandridae Goldfuss, 1820 (Amphibia, Urodela). Alytes, 26 (1–4), 1–85.

    Duméril, A. (1853) Prodrome de la classification des Reptiles Ophidiens. Typographie de Firmin Didot Frères, Paris, 139 + (1), Pl. 1–2.

    Duméril, A.M.C., Bibron, G. & Duméril, A.H.A. (1854) Erpétologie générale ou histoire naturelle complète des reptiles. Tome septième. Deuxième partie, comprenant l'histoire des serpents venimeux, Librairie Encyclopédique de Roret, Paris, i–xii + 781–1536, Pl. 71–84.

    Duvernoy, G.L. (1832) Mémoire sur les caractères tirés de l’anatomie pour distinguer les serpens venimeux des serpens non venimeux. Annales des Sciences naturelles, Paris, 26, 113–160, Pl. 5–10.

    Elera, C. de (1895) Catálogo sistemático de toda la fauna de Filipinas conocida hasta el presente, y á la vez el de la colección zoológica del museo de PP. Domenicos del Colegio-Universidad de Sto. Tomás de Manila. Volumen I. Vertebrados, Colegio de Santo Tomás, Manila, viii + (1) + 701 pp.

    Feuer, R.C. & Smith, H.M. (1972) The contributions of the 1822 works of Jarocki and Fleming to herpetological nomenclature. Great Basin Naturalist, 32 (1), 55–60.

    Fischer, G. (1813) Zoognosia. Tabulis synopticis illustrata, in usum prælectionum Academiae Imperialis Medico-chirurgicæ Mosquensis edita, auctore Gotthelf Fischer. Editio tertia, Classium, Ordinum, Generum illustratione perpetua aucta. Volumen primum. Tabulas synopticas generales et comparatives, nec non characterum quorundam explicationens iconographicam continens. Typis Nicolai Sergeidis Vsevolozsky, Mosquae [Moscow], xiv + 464 (+ 1 unnumbered page).

    Fitzinger, L.J.F.J. (1826) Neue Classification der Reptilien nach ihren natürlichen Verwandtschaften, nebst einer Verwandtschafts-Tafel und einem Verzeichnisse der Reptilien-Sammlung des k. k. zoolog. Museums zu Wien. J. G. Heubner, Wien, 8 + 66 pp.

    Fitzinger, L.J. (1843) Systema reptilium. Fasciculus primus. Amblyglossae. Braumüller & Seidel Bibliopolas, Vindobonae [Vienna], 106 + ix pp.

    Fitzinger, L.J. (1861) Die Ausbeute der österreichischen Naturforscher an Säugethieren und Reptilien während der Weltumsegelung Sr. Majestät Fregatte Novara. Sitzungsberichte der Mathematisch-Naturwissenschftlichen Classe der Kaiserlichen Akademie der Wissenschafte, 42, 383–416.

    Fleming, J. (1822) The philosophy of zoology; or a general view of the structure, functions, and classification of animals. Vol. II. Archibald Constable & Co, Edinburgh, 618 pp.

    Freycinet, L. (1815) Voyage de découvertes aux Terres Australes exécuté sur les corvettes Le Géographe, Le Naturaliste, et la goëlette Le Casuarina, pendant les années 1800, 1801, 1802, 1803 et 1804, sous le commandement du Capitaine de vaisseau N. Baudin. Navigation et Géographie. Publié par ordre de son Excellence le ministre de la Marine et des colonies. Avec un atlas. Imprimerie Royale, Paris, xvi + 591 pp.

    Giannasi, N., Thorpe, R.S. & Malhotra, A. (2001) The use of amplified fragment length polymorphism in determining species trees at fine taxonomic levels: analysis of a medically important snake, Trimeresurus albolabris. Molecular Ecology, 10, 419–426.

    Golay, P., Smith, H.M., Broadley, D.G., Dixon, J.R., McCarthy, C.J., Rage, J.-C., Schätti, B. & Toriba, M. (1993) Endoglyphs and other major venomous snakes of the world. A checklist. Azemiops S.A. Herpetological Data Center, Aïre–Genève, xv + 478 pp.

    Goldfuss, G. A. 1820. Handbuch der Zoologie. Zweite Abteilung. Systematische Übersicht der Gattungen. In. G. H. von Schubert, (Ed.), Handbuch der Naturgeschichte zum Gebrauche bei Vorlesungen, Dritter Theil, Zweite Abteilung, Johann Leonhard Schrag, Nürnberg, xxiv + 510 pp.

    Gray, J.E. (1825) A synopsis of the genera of Reptiles and Amphibia, with a description of some new species. Annals of Philosophy, London, (2) 10, 193–217.

    Gray, J.E. (1831) A synopsis of the species of the Class Reptilia. In: Griffith, E. & Pidgeon, E. (Ed.) The Animal Kingdom arranged in conformity with its organization, by the Baron Cuvier, member of the Institute of France, with additional descriptions of all the species hitherto named, and of many not before noticed. Volume the Ninth. The class Reptilia arranged by the Baron Cuvier, with specific descriptions. Appendix. Whittaker Treacher, and Co. London, 386 pp. + 110 pp. (Appendix), Pl. 1–55.

    Gray, J.E. (1840) Synopsis of the contents of the British Museum. 42nd edition. G. Woodfall, London, 370 pp.

    Gray, J.E. (1842) Synopsis of the species of rattle-snakes, or family of Crotalidae. Zoological Miscellany, 2, 47–51.

    Gray, J.E. (1849) Catalogue of the specimens of snakes in the collection of the British Museum. British Museum (Natural History), London, xv + 125 pp.

    Gray, J.E. (1853) Descriptions of some undescribed species of Reptiles collected by Dr. Joseph Hooker in the Khassia Mountains, East Bengal, and Sikkim Himalaya. Annals and Magazine of natural History, (2), 12 (59), 386–392.

    Guérin-Méneville, F.E. (1844) Iconographie du Règne Animal de G. Cuvier, ou Représentation d’après nature de l’une des espèces les plus remarquables et souvent non encore figurées, de chaque genre d’animaux. Avec un texte descriptif mis au courant de la science. Ouvrage pouvant servir d’atlas à tous les traités de zoologie. Tome III. Texte explicatif. Reptiles. J. B. Baillière, Paris, 1-23.

    Note. We follow here Zhao & Adler (1993: 372) for the date of publication of “Tome III” of this work.

    Guibé, J. & Roux-Estève, R. (1972) Les types de Schlegel (Ophidiens) présents dans les collections du Muséum National d’Histoire Naturelle de Paris. Zoologische Mededelingen, 47, 129–134.

    Gumprecht, A. (1997) Die Bambusottern der Gattung Trimeresurus Lacépède. Teil I: Die Chinesische Bambusotter Trimeresurus stejnegeri Schmidt, 1925. Sauria (Berlin), 19 (3), 9–30.

    Gumprecht, A. (1998) Die Bambusottern der Gattung Trimeresurus Lacépède Teil II: Die Großaugen - Bambusotter Trimeresurus macrops Kramer. Sauria (Berlin), 20 (3), 25–36.

    Gumprecht, A. (2007a) Asiatische Bambusottern. Terraria, 2 (2), 4–8.

    Gumprecht, A. (2007b) Zur aktuellen Taxonomie und Systematik der Asiatischen Bambusottern. Terraria, 2 (2), 9–12.

    Gumprecht, A. & Ryabov, S. (2002) Die Gattung Trimeresurus Lacépède, 1804. Zum Kenntnisstand der Forschung. Draco, 12, 31–44.

    Gumprecht A., Tillack F., Orlov N., Captain A. & Ryabov S. (2004) Asian Pitvipers. Geitje Books, Berlin, 368 pp.

    Günther, A.C.L.G. (1858) Catalogue of colubrine snakes in the collection of the British Museum. British Museum of Natural History, London, xvi + 281 pp.

    Günther, A.C.L.G. (1860a) On the Reptiles of Siam. Proceedings of the zoological Society, London, 28, 113–117, Pl. 23.

    Günther, A.C.L.G. (1860b) Contributions to a knowledge of the Reptiles of the Himalaya Mountains. Proceedings of the zoological Society, London, 28, 148–175, Pl. 25–28.

    Günther, A.C.L.G. (1864) The Reptiles of British India. Ray Society, London, xxvii + 452 pp., Pl. 1–26.

    Guo, P. & Wang, Y. (2011) A new genus and species of cryptic Asian green pitviper (Serpentes: Viperidae: Crotalinae) from southwest China. Zootaxa, 2918, 1–14.

    Hagen, B. (1890) Die Pflanzen- und Thierwelt von Deli auf der Ostküste Sumatras. Tijschr. kon. ned. Aadrijksk. Genootsch., Amsterdam, 2, 1–240.

    Harding, K.A. & Welch, K.R.G. (1980) Venomous snakes of the world. A checklist. Pergamon Press, Oxford, x + 188 pp.

    Haworth, A.H. (1825) A binary arrangement of the class Amphibia. The philosophical Magazine and Journal, London, 65, 372–373.

    Hemming, F. (Ed.) (1954) Opinion 260. Rejection for nomenclatorial purposes of the work by Meuschen (F.C.) issued in 1778 under the title Museum Gronovianum. Opinions and Declarations rendered by the International Commission on Zoological Nomenclature, 5 (21), 265–280, pl. 1–2.

    Hoge, A.R. & Romano Hoge, S.A.R.W.L. (1981) Poisonous snakes of the world. Part I. Check-list of the pit vipers Viperoidea, Crotalinae. Memórias do Instituto Butantan, 42/43 [1978/1979], 179–310.

    Iskandar, D.T. & Colijn, E. (2001) A checklist of Southeast Asian and New Guinean reptiles. Part I: Serpentes. Biodiversity Conservation Project (Indonesian Institute of Sciences–Japan International Cooperation Agency–The Ministry of Forestry), The Gibbon Foundation and Institut of Technology, Bandung, 195 pp.

    Jan, G. (1859) Plan d’une Iconographie descriptive des Ophidiens, et Description sommaire de nouvelles espèces de Serpents. Revue et Magasin de Zoologie pure et appliquée, (2), 11, 122–130.

    Jan, G. (1863) Elenco sistematico degli Ofidi descritti e designati per l’Iconografia generale, Tipografia di A. Lombardi, Milano, vii + 143 pp.

    Jangoux, M. (2005) L’expédition du capitaine Baudin aux Terres australes: les observations zoologiques de François Péron pendant la Première campagne (1801-1802). Annales du Muséum du Havre, 73, 1–35.

    Jarocki, F.P. (1822) Zoologia czyli Zwierzetopismo ogolne podlug Najnowszego Systematu ulozone przez Feliza Pawla Jarockiego. Tom 3. Gady i Plazy. Latkiewicza, Warsaw, [6 unnumbered pages], 184 pp., [11 unnumbered pages], Pl. 1–3.

    Klemmer, K. (1963) Liste der rezenten Giftschlangen. Elapidae, Hydrophiidae, Viperidae und Crotalidae. Behringwerk-Mitteilungen, Sonderband, Die Giftschlangen der Erde, Marburg / Lahn, 253–464.

    Kopstein, F. (1938) Herpetologische Notizen XVIII. Über die systematische Stellung der Art Trimeresurus gramineus (Shaw) von den Sunda-Inseln. Treubia, 16 (3), 325–333.

    Kramer, E. (1977) Zur Schlangenfauna Nepals. Revue suisse de Zoologie, 84 (3), 721–761.

    Kramer, E. (1978) Typenkatalog der Schlangen des NMB, Stand 1977. Revue suisse de Zoologie, 85 (3), 657–665.

    Kuch, U. (2002) Records of Trimeresurus insularis Kramer, 1977 from Bali, Indonesia. Herpetozoa, 15 (3/4), 180–182.

    Kuhl, H. (1824) Sur les reptiles de Java. Extrait d’une lettre adressée de Java en Hollande, par M. Kuhl, datée de Pjihorjavor au pied du Pangerango, le 18 juillet 1821 (Algem. Konst en Letterbode; 1822). Bulletin des Sciences naturelles et de Géologie (Section II du Bulletin Universel des Sciences naturelles), Paris, 2 (1), 79–83.

    Kuhl, H. & Van Hasselt, J.C. (1822) Uittreksels uit brieven van de Heeren Kuhl en van Hasselt, aan de Heeren C. J. Temminck, Th. Van Swinderen en W. de Haan. Algemeene Konst en Letterbode, 1 (7), 99–104.

    Lacépède, B.G.E.L. (Comte de) (1804) Mémoire sur plusieurs animaux de la Nouvelle Hollande dont la description n’a pas encore été publiée. Annales du Muséum d’Histoire naturelle, Paris, 4, 184–211, Pl. 55–58.

    Lesson, R.-P. (1829–1831) Observations générales sur les reptiles recueillis dans le voyage de la corvette La Coquille. In: Duperrey, L.I. (Ed.), Voyage autour du monde, exécuté par ordre du Roi, sur la corvette de sa Majesté, La Coquille, pendant les années 1822, 1823, 1824, et 1825, Zoologie, Vol. II, A. Bertrand, Paris, 1–65 + 465, Pl. 1-7.

    Note. We follow here Zhao & Adler (1993: 381) for the dates of publication of the parts of this work: 1829, Plates 1, 3, 5–7; 1830, Plates 2 & 4, pages 1–24; 1831, Pages 25–65 & 465.

    Leviton, A.E. (1964) Contributions to a review of Philippine snakes. V. The snakes of the genus Trimeresurus. Philippine Journal of Science, 93 (2), 251–276

    Leviton, A.E. (1968) The venomous terrestrial snakes of East Asia, India, Malaya, and Indonesia. In: W. Bücherl, E.E. Buckley & V. Deulofeu (Ed.), Venomous animals and their venoms, Volume I, Venomous vertebrates, Academic Press, New York & London, 529–576.

    Lichtenstein, M.H.C. (1823) Verzeichniss der Doubletten des zoologischen Museums der Königl. Universität zu Berlin nebst Beschreibung vieler bisher unbekannten Arten von Säugethieren, Vögeln, Amphibien und Fischen. T. Trautwein, Berlin, x + 118 pp.

    Lichtenstein, H., Weinland, D. & Martens, E. von (1856) Nomenclator reptilium et amphibiorum Musei Zoologici Berolinensis. Namenverzeichniss der in der zoologischen Sammlung der Königlichen Universität zu Berlin aufgestellten Arten von Reptilien und Amphibien nach ihren Ordnungen, Familien und Gattungen. Königlichen Akademie der Wissenschaften, Berlin, iv + 48 pp.

    Mahendra, B.C. (1984) Handbook of the snakes of India, Ceylon, Burma, Bangladesh and Pakistan. Annals of Zoology, Agra, (B), 22, i–xvi + 1–412.

    Maki, M. (1931) A monograph of the snakes in Japan. Dai-Ichi Shobo, Tokyo, Vol. I, (8 unnumbered pages) + 240 pp. [English text].

    Malhotra, A. & Thorpe, R.S. (2004a) A phylogeny of four mitochondrial gene regions suggests a revised taxonomy for Asian pitvipers (Trimeresurus and Ovophis). Molecular Phylogenetics and Evolution, 32 (1), 83–100.

    Malhotra, A. & Thorpe, R.S. (2004b) Reassessment of the validity and diagnosis of the pitviper Trimeresurus venustus Vogel, 1991. Herpetological Journal, London, 14, 21–33.

    Malhotra, A. & Thorpe, R.S. (2004c) Maximizing information in systematic revisions: a combined molecular and morphological analysis of a cryptic green pitviper complex (Trimeresurus stejnegeri). Biological Journal of the Linnean Society, 82, 219–235.

    Malhotra, A., Thorpe, R.S., Mrinalini & Stuart, B. L. (2011) Two new species of pitviper of the genus Cryptelytrops Cope 1860 (Squamata: Viperidae: Crotalinae) from Southeast Asia. Zootaxa, 2757, 1–23.

    Maslin, T.P. (1942) Evidence for the separation of the crotalid genera Trimeresurus and Bothrops, with a key to the genus Trimeresurus. Copeia, 1942 (1), 18–24.

    McDiarmid, R.W., Campbell, J.A. & Touré, T’S.A. (1999) Snake species of the world. A taxonomic and geographical reference. Volume 1. The Herpetologists’ League, Washington DC, xi + 511 pp.

    McKay, J.L. (2006) A field guide to the Amphibians and Reptiles of Bali. Krieger Publishing Company, Malabar, Florida, vii + 138 pp.

    Mell, R. (1922) Beiträge zur Fauna sinica. I Die Vertebraten Südchinas; Feldlisten und Feldnoten der Säuger, Vögel, Reptilien, Batrachier. Archiv für Naturgeschichte, Berlin, (A), 80 (10), 1–134.

    Merrem, B. (1820) Versuch eines Systems der Amphibien. Johann Christian Krieger, Marburg, xvi + 191 pp., 1 plate.

    Mertens, R. (1930) Die Amphibien und Reptilien der Inseln Bali, Lombok, Sumbawa und Flores (Beiträge zur Fauna der Kleinen Sunda-Inseln, I). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 42 (3), 115–344, Pl. 1–9.

    Meuschen, F.C. (1778) Museum Gronovianum, sive Index Rerum Naturalium tam Mammalium, Amphibiorum, Piscium, Insectorum, Conchyliorum, Zoophytorum, Plantarum et Mineralium Exquisitissimorum quam Artefactorum nonnullorum. Inter quae eminet Herbarium siccum Plantarum a Tournefortio, Claytonio, Linnaeo aliisque Botanicis collectarum. Quae omnia multa et magnis Sumptibus sibi comparavit Vir Amplissimus et Celeberrimus Laur. Theod. Gronovius. Lugduni Batavorum [Leyden], Theodorus Haak, 1–252 pp.

    Mocquard, F. (1915) Les genres Trimeresurus et Lachesis ne sont pas identiques. Bulletin du Muséum d’Histoire naturelle, Paris, 21 (4), 115–117.

    Müller, S. & Schlegel, H. (1845) Over de, in den Indischen Archipel voorkomende soorten van het slangengeslacht Trigonocephalus. In: C. J. Temminck (Ed.), Verhandelingen over de natuurlijke geschiedenis der Nederlandsche overzeesche bezittingen, door de Leden der Natuurkundige Commissie in Ost-Indië en andere Schrijvers, Zoologie, N° 7, Reptilia, S. & J. Luchtmans & C. C. Van der Hoek, Leiden, 49–58, Pl. 7 (1842).

    Note. The text on pitvipers is dated 1845, but the corresponding plate was published in 1842 (Plate 7, description of Trigonocephalus formosus). The complete part on Reptilia includes 72 pages text and 10 plates. It is divided into several chapters, each bearing a different title, and variously written by Müller & Schlegel or Schlegel & Müller. According to Zhao & Adler (1993: 398), the dates of publication of this part are as follows: 1839, Pages 1–8, Plates 1–3 ; 1840, Plate 4 ; 1841, Pages 9–28, Plates 5-6; 1842, Plate 7; 1843, Plates 8–10 ; 1845, Pages 29–72.

    Neave, S.A. (Ed.) (1940) Nomenclator zoologicus. Vol. IV. Q–Z and supplement. Zoological Society of London, London, iv + 758 pp.

    Oken, L. (1836) Allgemeine Naturgeschichte für alle Stände. Sechster Band, oder Thierreich, Dritter Band. Hoffmann’sche Verlags-Buchhandlung, v + 698 pp.

    Oppel, M. (1811a) Suite du Ier. mémoire sur la classification des reptiles. Annales du Muséum d’Histoire naturelle, Paris, 16, 376–393.

    Oppel, M. (1811b) Die Ordnungen, Familien und Gattungen der Reptilien als Prodrom einer Naturgeschichte derselben. Joseph Lindauer, München, xii + 71 pp.

    Orlov, N., Ananjeva, N., Barabanov, A., Ryabov, S. & Khalikov, R. (2002a) Diversity of vipers (Azemiopinae, Crotalinae) in East, Southeast, and South Asia: annotated checklist and natural history data (Reptilia: Squamata: Serpentes: Viperidae). In: Fritz, U. (Ed.), Collectanea Herpetologica, Essays in honour of Fritz Jürgen Obst, Faunistische Abhandlungen Staatliches Museum für Tierkunde Dresden, 23 (10), 177–218.

    Orlov, N., Ananjeva, N. & Khalikov, R. (2002b) Natural history of Pitvipers in eastern and southeastern Asia. In: Schuett, G.W., Höggren, M., Douglas, M.E. & Greene, H.W. (Ed.), Biology of the Vipers, Eagle Mountain Publishing, Eagle Mountain, Utah, 345–359.

    O’Shea, M. (2005) Venomous snakes of the world. New Holland Publishers (UK) Ltd, London–Cape Town–Sidney–Auckland, 160 pp.

    Ouwens, P.A. (1916) De voornaamste giftslangen van Nederlandsch Oost-Indië. Met een verhandeling over de verschijnselen en over de behandeling van giftigen slangenbeet door Dr. G. W. Kieweit de Jonge. E. J. Brill, Leiden, 1–22, (1), Pl. 1–19.

    Peters, W.C.H. (1862) Über die craniologischen Verschiedenheiten der Grubenottern (Trigonocephali) und über eine neue Art der Gattung Bothriechis. Monatsberichte der königlich Akademie der Wissenschaften zu Berlin, 1862 (December), 670–674.

    Peters, W.C.H. (1872) Übersicht der von den Herren M.se G. Doria und D.r O. Beccari in Sarawack auf Borneo von 1865 bis 1868 gesammelten Amphibien. Annali del Museo Civico di Storia Naturale di Genova, (1), 3, 27–45, Pl. 2–6.

    Pope, C.H. & Pope, S.H. (1933) A study of the green pit-vipers of southeastern Asia and Malaysia, commonly identified as Trimeresurus gramineus (Shaw), with description of a new species from Peninsular India. American Museum Novitates, 620, 1–12.

    Raffles, T.S. (1822) Second part of the descriptive catalogue of a zoological collection made in the island of Sumatra and its vicinity. Transactions of the Linnean Society of London, 13 (2), 277–340.

    Rafinesque, C.S. (1815) Analyse de la nature ou Tableau de l’univers et des corps organisés. Jean Barravecchia, Palerme, 124, 1 plate.

    Regenass, U. & Kramer, E. (1981) Zur Systematik der grünen Grubenottern der Gattung Trimeresurus (Serpentes, Crotalidae). Revue suisse de Zoologie, 88 (1), 163–205.

    Rigondet, G. (2002) François Péron 1775-1810 et l’expédition du commandant Nicolas Baudin. Les Français à la découverte de l’Australie. Editions des Cahiers Bourbonnais, Charroux (Allier, France), 319 pp (+ 4 unnumbered pages).

    Ritgen, F. A. (1828) Versuch einer natürlichen Eintheilung der Amphibien. Verhandlungen der Kaiserlichen Leopoldinisch-Carolinischen Akademie der Naturforscher, 6, 247–284.

    Roux-Estève, R. (1979) Liste des Amphibiens et Reptiles des collections du Muséum National d’Histoire Naturelle de Paris, récoltés par Lesueur (1778–1846). Bulletin trimestriel de la Société de Géologie de Normandie et des Amis du Muséum du Havre, 66 (3), 25–29.

    Ruiz, J. (1952) Sobre a distincão genérica dos Crotalidae (Ophidia: Crotaloidea) baseada em alguns caracteres osteológicos (Nota preliminar). Memórias do Instituto Butantan, 23, 109–114.

    Rüppell, E. (1845) Verzeichniss der in dem Museum der Senckenbergischen naturforschenden Gesellschaft aufgestellten Sammlungen. Dritte Abtheilung: Amphibien. Verh. Mus. Senckenberg, 3, 295–316.

    Russell, P. (1796) An account of Indian Serpentes collected on the coast of Coromandel; containing descriptions and drawings of each species, together with experiments and remarks on their several poisons. Presented to the Hon. The Court of the East India Company, and published by their order, under the Superintendence of the author, George Nicol, London, viii + 91 pp., Pl. 1–46.

    Russell, P. (1801–1809) A continuation of an account of Indian Serpents; containing descriptions and figures from specimens and drawings, transmitted from various parts of India. Presented to the Hon. The Court of the East India Company, and published by their order, under the Superintendence of the author. G. & W. Nicol London, v + xv + 53 pp. (+ 4 unnumbered pages), Pl. 1–45.

    Note. According to Zhao & Adler (1993: 396), this book was published in five sections, as follows: Section 1, 1801: pages i–v, 1–12, plates 1–10; Section 2, 1802: pages 13–20, plates 11–18; Section 3, 1804: pages 21–28, plates 19–24; Section 4, 1807: pages 29–38, plates 25–32; Section 5, 1809 (in fact probably 1810): pages ix–xv, 39-53, (4), portrait, plates 33–45. See Zhao & Adler (1993) for additional information about this work.

    Sabrosky, C.W. (1999) Family-group names in Diptera. An annotated catalogue. Myia, 10, 1–576.

    Saint Girons, H. (1972) Les serpents du Cambodge. Mémoires du Muséum national d’Histoire naturelle, (Nouvelle série), (A, Zoologie), 74, 1–170, Pl. 1–42.

    Sanders, K.L., Malhotra, A. & Thorpe, R.S. (2004) Ecological diversification in a group of Indomalayan Pitvipers (Trimeresurus): convergence in taxonomically important traits has implications for species identification. Journal of evolutionary Biology, 17, 721–731.

    Schätti, B. & Perret, J.-L. (1997) Catalogue révisé des types d'amphibiens et de reptiles du Muséum d'histoire naturelle de Genève. Revue suisse de Zoologie, 104(2), 357–370.

    Schinz, H.R. (1833-1835) Naturgeschichte und Abbildungen der Reptilien. Nach den neuesten Systemen zum gemeinnützigen Gebrauche entworfen und mit Berücksichtigung für den Unterricht der Jugend bearbeitet. Nach der Natur und dem vorzüglichsten Originalien gezeichnet und lithographirt von K. J. Brodtmann. Des Thierreichs dritter Theil. Brodtman Lithographischer Anhalt, Shaffhausen, iv + 240, pp., Pl. 1–102.

    Note. A discussion on the dates of publication of this work, initially published in 17 fascicules, appeared in David & Ineich (1999). Pages 97–198 were published in 1834.

    Schlegel, H. (1826) Notice sur l’erpétologie de l’Ile de Java; par M. Boïé. (Ouvrage manuscrit). Bulletin universel des Sciences et de l’Industrie de Férussac (Section II du Bulletin des Sciences naturelles et de Géologie), 9 (2), 233–240.

    Schlegel, H. (1827) Erpetologische Nachrichten. Isis von Oken, 20 (3), col. 281–294.

    Schlegel, H. (1837) Essai sur la physionomie des serpens. II. Partie descriptive. J. Kips, H. Hz. & W. P. Van Stockhum, The Hague, 606 + xv pp. Atlas, (2 unnumbered pages), Pl. 1–21, 3 maps, 2 tables + (2 unnumbered pages).

    Sclater, W.L. (1891) List of snakes in the Indian Museum. The Indian Museum, Calcutta, x + 79 pp.

    Seba, A. (1735) Locupletissimi Rerum Naturalium Thesauri Accurata Descriptio, et Iconibus Artificiosissimus Expressio, per Universam Physices Historiam. Opus, cui, in hoc Rerum Genere, Nullum Par Existit. Ex Toto Terrarum Orbe Collegit, Digessit, Descripsit, et Depingendum Curavit. Tomus II. Janssonio-Waesbergios, J. Wetstenium & Gul. Smith Amstelaedami [Amsterdam], (2 unnumbered pages) + (30 unnumbered pages) + 154 pp., Pl. 1–114.

    Shaw, G. (1802) General zoology, or systematic natural history. Vol. III. Part II. Amphibia. G. Kearsley, London, i–vii + 313–615, Pl. 87–140.

    [Reprinted in 1999 by the Society for the Study of Amphibians and Reptiles, Ithaca, New York].

    Simpson, G.G. (1940) Types in modern taxonomy. American Journal of Science, 238, 413–431.

    Smith, M.A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo-chinese subregion. Reptilia and Amphibia. Vol. III, Serpentes. Taylor & Francis, London, xii + 583.

    Stejneger, L.H. (1907) Herpetology of Japan and adjacent territory. Bulletin of the United States national Museum, 58, i–xx + 1–577, Pl. 1–35.

    Stejneger, L.H. (1927) The Green pit viper, Trimeresurus gramineus, in China. Proceedings of the United States national Museum, 72 (19), 1–10.

    Stoliczka, F. (1870) Observations on some Indian and Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal, 39 (2), 134–228, Pl. 8–12.

    Swainson, W. (1839) On the natural history and classification of Fishes, Amphibians, and Reptiles. Vol. II. Longman, Orme, Brown, Green, & Longmans, and John Taylor, London, vi + 452 pp.

    Takara, T. (1962) Studies on terrestrial snakes in the Ryukyu Archipelago. The Science Bulletin of the Division of Agriculture, Home Economics and Engineering, University of the Ryukyus, 9 (1), 1–202, Pl. 1–22. (In Japanese, English summary).

    Taylor, E.H. (1965) The serpents of Thailand and adjacent waters. University of Kansas Science Bulletin, 45 (9), 609–1096.

    Theobald, W. (1868) Catalogue of Reptiles in the Museum of the Asiatic Society of Bengal, 1866. Journal of the Asiatic Society of Bengal, 88 (37), 1–88 + i–iii, Pl. 1–4.

    Theobald, W. (1876) Descriptive catalogue of the Reptiles of British India. Thacker, Spink & Co., Calcutta, x + 238 + xxxviii + xiii, pp., Pl. 1.

    Traill, T.S. (1843) Essay of the physiognomy of Serpents. By H. Schlegel, doctor in Philosophy, Conservator of the Museum of the Netherlands, member of several learned societies. Translated by Thos. Stewart Traill, M.D., F.R.S.E., Regius Professor of Medical jurisprudence in the University of Edinburgh, &c. &c., Maclachlan, Stewart & Company, Edinburgh, vii + 254 pp., Corrigenda, Pl. 1–2.

    Ulber, T.M. (Ed.) (1993) Boie, 1827. Bemerkungen über Merrem’s Versuch eines Systems der Amphibien, Marburg. 1820. Erste Lieferung: Ophidier. / Remarks on Merrem’s Attempt of a System of the Amphibians, Marburg, by Friedrich Boie of Kiel. 1st instalment. Ophidians. Herprint International, Bredell (South Africa), Appendix, 323–380.

    Vogel, G. (2006) Venomous snakes of Asia - Giftschlangen Asiens. Edition Chimaira, Frankfurt am Main & Aqualog Verlag ACS, Rodgau, 148 pp.

    Wagler, J.G. (1830) Natürliches System der Amphibien, mit vorangehender Classification der Säugthiere und Vögel. Ein Beitrag zur vergleichenden Zoologie, J.G. Cotta Buchhandlung München, vi + 354 pp.

    Wagler, J.G. (1833) Deutung der in Seba’s Thesauro rerum naturalium T. 1. et 2. enthaltenen Abbildungen von Lurchen, mit critischen Bemerkungen. Isis von Oken, 26 (9), col. 885–905.

    Wallach, V. & Pauwels, O.S.G. (2008) The systematic status of Cathetorhinus melanocephalus Duméril & Bibron, 1844 (Serpentes: Typhlopidae). Hamadryad, 33, 39–47.

    Wallach, V., Wüster, W. & Broadley, D.G. (2009). In praise of subgenera: taxonomic status of cobras of the genus Naja Laurenti (Serpentes: Elapidae). Zootaxa, 2236, 26–36.

    Welch, K.R.G. (1988) Snakes of the Orient: a checklist. Robert F. Krieger Publishing Co., Malabar (Florida), vii + 183 pp.

    Werner, F. (1922) Synopsis der Schlangenfamilien der Amblycephaliden und Viperiden nebst Übersicht über die kleineren Familien und die Colubriden der Acrochordinen-Gruppe. Auf Grund des Boulenger’schen Schlengenkataloges (1893-1896). Archiv für Naturgeschichte, Berlin, (A), 88 (8), 185–244.

    Williams, K.L. & Wallach, V. (1989) Snakes of the world. Vol. I. Synopsis of snake generic names. Robert E. Krieger, Malabar (Florida), viii + 234 pp.

    Zhao, E. & Adler, K. (1993) Herpetology of China. Contribution to Herpetology, 10, Society for the Study of Amphibians and Reptiles, Athens (Ohio), 522 pp. + Pl. 1–48.

    Ziegler, T. (2002) Die Amphibien und Reptilien eines Tieflandfeuchtwald-Schutzgebietes in Vietnam. Natur und Tier-Verlag GmbH, Münster, 342 pp.