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Type: Articles
Published: 2012-01-24
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On a collection of hydroids (Cnidaria, Hydrozoa, Hydroidolina) from the west coast of Sweden, with a checklist of species from the region

Department of Natural History, Royal Ontario Museum, 100 Queen’s Park, Toronto, Ontario, Canada M5S 2C6
Cnidaria Anthoathecata Kattegat Kosterhavet Leptothecata marine invertebrates northwest Europe Skagerrak taxonomy zoological nomenclature

Abstract

An account is given of 48 species identified in a small collection of hydroids from the west coast of Sweden. Assigned to39 genera and 18 families, most are well-known from boreal waters of the North Atlantic Ocean. Campanulina paniculaG.O. Sars, 1874 is referred to Racemoramus, a new genus of leptothecates distinguished in part by having monopodialgrowth, rigid and non-segmented hydrocauli, and grape-like clusters of hydrothecae that arise at irregular intervals fromthe stem. The name Sertularia dumosa (now Lafoea dumosa) is taken to have been made available by Fleming (1820), notFleming (1828). As such, it is not threatened by a virtually unused subjective synonym (Lafoea cornuta Lamouroux,1821). Aselomaris michaeli Berrill, 1948 and Rhizorhagium navis Millard, 1959 are regarded as synonyms and combinedunder the binomen Pachycordyle michaeli (Berrill, 1948). Confusion in the literature over the identity of Sertularia teneraG.O. Sars, 1874 is addressed, and distinctive characters of the species are discussed. A species often misidentified as S.tenera is assigned to Sertularia spitzbergensis (Jäderholm, 1909); the name spitzbergensis is elevated in rank from sub-specific to specific, and assigned priority over its simultaneous synonym siberica Jäderholm, 1909 under the First ReviserPrinciple. Gonothecae of Clytia gigantea (Hincks, 1866) (type locality: Lamlash Bay, UK) are reported for the first timein material from Europe. Although much like Clytia hemisphaerica, this species is retained as valid due to its remarkablyrobust colony form, elongate and gradually tapered hydrothecae, and linguiform rather than wavy or pointed marginalcusps. Eudendrium album Nutting, 1896, Pachycordyle michaeli (Berrill, 1948), and the hydroid stage of Halitholus cir-ratus Hartlaub, 1913 are reported for the first time from the study area. A checklist of hydroids from the study area is included.

References

  1. Agassiz, A. (1865) Illustrated catalogue of the Museum of Comparative Zoölogy, at Harvard College. No. II. North American Acalephae. Sever & Francis, Cambridge, Massachusetts, 234 pp.

    Agassiz, E.C. & Agassiz, A. (1865) Seaside studies in natural history. Marine animals of Massachusetts Bay. Radiates. Ticknor & Fields, Boston, 155 pp.

    Agassiz, L. (1862) Contributions to the natural history of the United States of America. Vol. IV. Little, Brown & Company, Boston, 380 pp.

    Alder, J. (1856) A notice of some new genera and species of British hydroid zoophytes. Annals and Magazine of Natural History, series 2, 18, 353–362.

    Alder, J. (1857) A catalogue of the zoophytes of Northumberland and Durham. Transactions of the Tyneside Naturalists’ Field Club, 3, 93–162.

    Alder, J. (1859) Descriptions of three new species of sertularian zoophytes. Annals and Magazine of Natural History, series 3, 3, 353–356.

    Alder, J. (1860) Descriptions of a zoophyte and two species of Echinodermata new to Britain. Annals and Magazine of Natural History, series 3, 5, 73–75.

    Alder, J. (1862) Descriptions of some new and rare zoophytes found on the coast of Northumberland. Annals and Magazine of Natural History, series 3, 9, 311–317.

    Allman, G.J. (1859) Notes on the hydroid zoophytes. I. Laomedea flexuosa, Hincks. II. The extra-capsular medusiform sporosacs (“meconidia”) of Laomedea, and the determination of the species in which they are found. III. Coryne eximia, nov. sp., and its medusa. Annals and Magazine of Natural History, series 3, 4, 137–144.

    Allman, G.J. (1863) Notes on the Hydroida. I.–On the structure of Corymorpha nutans. II. Diagnoses of new species of Tubularidae obtained, during the autumn of 1862, on the coasts of Shetland and Devonshire. Annals and Magazine of Natural History, series 3, 11, 1–12.

    Allman, G.J. (1864) On the construction and limitation of genera among the Hydroida. Annals and Magazine of Natural History, series 3, 13, 345–380.

    Allman, G.J. (1872) A monograph of the gymnoblastic or tubularian hydroids.Conclusion of Part I, and Part II, containing descriptions of the genera and species of the Gymnoblastea. Ray Society, London, pp. 155–450.

    Allman, G. J. (1874a) Report on the Hydroida collected during the expeditions of H.M.S. ‘Porcupine.’ Transactions of the Zoological Society of London, 8, 469–481.

    Allman, G.J. (1874b) On the diagnosis of new genera and species of hydroids. Nature, 11, 179.

    Allman, G.J. (1877) Report on the Hydroida collected during the exploration of the Gulf Stream by L. F. de Pourtalès, assistant, United States Coast Survey. Memoirs of the Museum of Comparative Zoölogy at Harvard College, 5(2), 1–66.

    Allman, G.J. (1883) Report on the Hydroida dredged by H.M.S. Challenger during the years 1873–76. Part I.–Plumularidae. Report on the Scientific Results of the Voyage of H.M.S. Challenger during the Years 1873–76, Zoology, 7(20), 1–54.

    Allman, G.J. (1888) Report on the Hydroida dredged by H.M.S. Challenger during the years 1873–76. Part II—The Tubularinae, Corymorphinae, Campanularinae, Sertularinae, and Thalamophora. Report on the Scientific Results of the Voyage of H.M.S. Challenger during the Years 1873–76, Zoology, 7(23), 1–90.

    Altuna, A. (2007) Bathymetric distribution patterns and biodiversity of benthic Medusozoa (Cnidaria) in the Bay of Biscay (north-eastern Atlantic). Journal of the Marine Biological Association of the United Kingdom, 87, 681–694.

    Anger, K. (1972) Dipurena spongicola sp. n. (HYDROZOA, CORYNIDAE), ein in Schwämmen lebender Hydroidpolyp aus dem Kattegat und der nördlichen Kieler Bucht. Kieler Meeresforschungen, 28, 80–83.

    Antsulevich, A.E. (1987) Gidroidy shel'fa Kuril'skykh ostrovov. Hydroids from the shelf waters of Kurile Islands. Zoologicheskiï Institut, Akademiya Nauk SSSR, 1–165.

    Babić, K. (1904) Uebersicht der Hydroidpolypen des adriatischen Meeres. Glasnik Hrvatskoga Naravoslovnoga Društva, Godina, 15, 201–220.

    Bandel, K. & Wedler, E. (1987) Hydroid, amphineuran and gastropod zonation in the littoral of the Caribbean Sea, Colombia. Senckenbergiana Maritima, 19, 1–129.

    Bedot, M. (1916) Sur le genre Kirchenpaueria. Revue Suisse de Zoologie, 24, 637–648.

    Berghahn, R. & Offermann, U. (1999) Laboratory investigations on larval development, motility and settement of white weed (Sertularia cupressina L.) – in view of its assumed decrease in the Wadden Sea. Hydrobiologia, 392, 233–239.

    Berrill, N.J. (1948) The life cycle of Aselomaris michaeli, a new gymnoblastic hydroid. Biological Bulletin, 95, 289–295.

    Berrill, N.J. (1949) Growth and form in gymnoblastic hydroids. I. Polymorphic development in Bougainvillia and Aselomaris. Journal of Morphology, 84, 1–30.

    Billard, A. (1922) Note sur deux espèces d’hydroïdes du littoral d’Ostende. Annales de la Société Royale Zoologique et Malacologique de Belgique, 52, 135–139.

    Birula, A. (1897) Materialui dlya biologii i zoogeografii preimushchestvenno russkikh morei. II. Hydrozoa, Polychaeta i Crustacea, sobrannije d-rom A.S. Botkinim v Eniseyskoy i Obskoy gubach, letom 1895 goda. Annuaire du Musée Zoologique de l’Académie Impériale des Sciences de St.-Pétersbourg, 2, 78–116.

    Bonnevie, K. (1898) Zur Systematik der Hydroiden. Zeitschrift für Wissenschaftliche Zoologie, 63, 465–495.

    Bonnevie, K. (1899) Hydroida. Den Norske Nordhavs-Expedition 1876-1878, Zoologi, 7 (26), 1–103.

    Bouillon, J., Medel, M.D., Pagès, F., Gili, J.-M., Boero, F. & Gravili, C. (2004) Fauna of the Mediterranean Hydrozoa. Scientia Marina, 68 (Suppl. 2), 5–438.

    Bouillon, J., Gravili, C., Pagès, F., Gili, J.-M. & Boero, F. (2006) An introduction to Hydrozoa. Mémoires du Muséum National d’Histoire Naturelle, 194, 591 pp.

    Brinckmann, A. (1965) The life cycle of the medusa Cirrholovenia tetranema Kramp 1959 (Leptomedusae, Lovenellidae) with a hydroid of the genus Cuspidella Hincks. Canadian Journal of Zoology, 43, 13–15.

    Brinckmann-Voss, A. (1996) Seasonality of hydroids (Hydrozoa, Cnidaria) from an intertidal pool and adjacent subtidal habitats at Race Rocks, off Vancouver Island, Canada. Scientia Marina, 60, 89–97.

    Broch, H. (1903) Die von dem norwegischen Fischereidampfer „Michael Sars” in den Jahren 1900–1902 in dem Nordmeer gesammelten Hydroiden. Bergens Museums Aarbog, 1903 (9), 1–14.

    Broch, H. (1908) Hydroiduntersuchungen I. Tecaphore Hydroiden von dem nördlichen Norwegen nebst Bemerkungen über die Variation und Artbegrenzung der nordischen Lafoëa-Arten. Tromsø Museums Aarshefter, 29, 27–40.

    Broch, H. (1910) Die Hydroiden der Arktischen Meere. Fauna Arctica, 5, 127–248.

    Broch, H. (1916) Hydroida (Part I.). The Danish Ingolf Expedition, 5(6), 66 pp.

    Broch, H. (1918) Hydroida (Part II.). The Danish Ingolf Expedition, 5(7), 205 pp.

    Broch, H. (1928) Hydrozoa I. In: Grimpe, G. & Wagler, E. (Eds.), Die Tierwelt der Nord-und Ostsee, 3.b, 1–100.

    Browne, E.T. (1902) A preliminary report on hydromedusae from the Falkland Islands. Annals and Magazine of Natural History, series 7, 9, 272–284.

    Cairns, S.D., Calder, D.R., Brinckmann-Voss, A., Castro, C.B., Fautin, D.G., Pugh, P.R., Mills, C.E., Jaap, W.C., Arai, M.N., Haddock, S.H.D., & Opresko, D.M. (2002) Common and scientific names of aquatic invertebrates from the United States and Canada: Cnidaria and Ctenophora. Second Edition. American Fisheries Society Special Publication, 28, 115 pp.

    Calder, D.R. (1970) Thecate hydroids from the shelf waters of northern Canada. Journal of the Fisheries Research Board of Canada, 27, 1501–1547.

    Calder, D.R. (1971) Hydroids and hydromedusae of southern Chesapeake Bay. Virginia Institute of Marine Science, Special Papers in Marine Science, 1, 1–125.

    Calder, D.R. (1972) Some athecate hydroids from the shelf waters of northern Canada. Journal of the Fisheries Research Board of Canada, 29, 217–228.

    Calder, D.R. (1974) The mud-dwelling hydrozoan Boreohydra simplex in the western North Atlantic. Journal of the Fisheries Research Board of Canada, 31, 1666–1667.

    Calder, D.R. (1975) Biotic Census of Cape Cod Bay: hydroids. Biological Bulletin, 149, 287–315.

    Calder, D.R. (1988) Shallow-water hydroids of Bermuda. The Athecatae. Royal Ontario Museum, Life Sciences Contributions, 148, 1–107.

    Calder, D.R. (1990) Seasonal cycles of activity and inactivity in some hydroids from Virginia and South Carolina, U.S.A. Canadian Journal of Zoology, 68, 442–450.

    Calder, D.R. (1991) Shallow-water hydroids of Bermuda. The Thecatae, exclusive of Plumularioidea. Royal Ontario Museum, Life Sciences Contributions, 154, 1–140.

    Calder, D.R. (2004) Subtidal hydroids (Cnidaria) of Northumberland Strait, Atlantic Canada, with observations on their life cycles and distributions. Canadian Field-Naturalist, 117, 555–564. (Although dated 2003, this article did not appear until 2004).

    Calder, D.R. (2010) Some anthoathecate hydroids and limnopolyps (Cnidaria, Hydrozoa) from the Hawaiian archipelago. Zootaxa, 2590, 1–91.

    Calder, D.R. & Cairns, S.D. (2009) Hydroids (Cnidaria: Hydrozoa) of the Gulf of Mexico. In: Felder, D.L. & Camp, D.K. (Eds.), Gulf of Mexico. Origin, waters, and biota. Volume 1, Biodiversity. Texas A&M University Press, College Station, pp. 381–394.

    Calder, D.R. & Vervoort, W. (1986) Plicatotheca anitae, a new genus and species of thecate hydroid from Bermuda and South Africa. Canadian Journal of Zoology, 64, 2021–2023.

    Calder, D.R., Vervoort, W. & Hochberg, F.G. (2009) Lectotype designations of new species of hydroids (Cnidaria, Hydrozoa), described by C. M. Fraser, from Allan Hancock Pacific and Caribbean Sea Expeditions. Zoologische Mededelingen, 83, 919–1058.

    Christiansen, B.O. (1972) The hydroid fauna of the Oslo Fiord in Norway. Norwegian Journal of Zoology, 20, 279–310.

    Ciamician, J. (1880) Ueber Lafoea parasitica n.sp. Zeitschrift für Wissenschaftliche Zoologie, 33, 673–676.

    Clark, S.F. (1875) Descriptions of new and rare species of hydroids from the New England coast. Transactions of the Connecticut Academy of Arts and Sciences, 3, 58–66.

    Clark, S.F. (1877) Report on the hydroids collected on the coast of Alaska and the Aleutian Islands, by W.H. Dall, U.S. Coast Survey, and party, from 1871 to 1874 inclusive. Proceedings of the Academy of Natural Sciences of Philadelphia, 1876, 28, 209–238.

    Clarke, S.F. (1888) Branch III.–Coelenterata. Class I.–Hydrozoa. Order I.–Hydroidea. In: Kingsley, J.S. (Ed.), The Riverside Natural History. Volume I. Lower invertebrates. Kegan Paul, Trench & Co., London, pp. 73–89.

    Clarke, S.F. (1907) Reports on the scientific results of the expedition to the eastern tropical Pacific, in charge of Alexander Agassiz, by the U.S. Fish Commission steamer “Albatross,” from October, 1904, to March, 1905, Lieut.-Commander L. M. Garrett, U.S.N., commanding. VIII. The hydroids. Memoirs of the Museum of Comparative Zoölogy at Harvard College, 35(1), 1–18.

    Cockerell, T.D.A. (1911) The nomenclature of the hydromedusae. Proceedings of the Biological Society of Washington, 24, 77–86.

    Collins, A.G. (2000) Towards understanding the phylogenetic history of Hydrozoa: hypothesis testing with 18S gene sequence data. Scientia Marina, 64 (Suplemento 1), 5–22.

    Cornelius, P.F.S. (1975a) The hydroid species of Obelia (Coelenterata, Hydrozoa: Campanulariidae), with notes on the medusa stage. Bulletin of the British Museum (Natural History), Zoology, 28, 249–293.

    Cornelius, P.F.S. (1975b) A revision of the species of Lafoeidae and Haleciidae (Coelenterata: Hydroida) recorded from Britain and nearby seas. Bulletin of the British Museum (Natural History), Zoology, 28, 375–426.

    Cornelius, P.F.S. (1977) On the nomenclature of the hydroid, Candelabrum phrygium (Fabricius, 1780) (=Myriothela phrygia, Arum cocksi). Journal of the Marine Biological Association of the United Kingdom, 57, 521–524.

    Cornelius, P.F.S. (1978) The genus names Calicella Hincks and Calycella Hincks (Coelenterata: Hydrozoa). Bulletin of the British Museum (Natural History), Zoology, 33, 233–234.

    Cornelius, P.F.S. (1979) A revision of the species of Sertulariidae (Coelenterata: Hydroida) recorded from Britain and nearby seas. Bulletin of the British Museum (Natural History), Zoology, 34, 243–321.

    Cornelius, P.F.S. (1982) Hydroids and medusae of the family Campanulariidae recorded from the eastern North Atlantic, with a world synopsis of genera. Bulletin of the British Museum (Natural History), Zoology, 42, 37–148.

    Cornelius, P.F.S. (1990) European Obelia (Cnidaria, Hydroida): systematics and identification. Journal of Natural History, 24, 535–578.

    Cornelius, P.F.S. (1992a) Medusa loss in leptolid Hydrozoa (Cnidaria), hydroid rafting, and abbreviated life-cycles among their remote-island faunae: an interim review. Scientia Marina, 56, 245–261.

    Cornelius, P.F.S. (1992b) The Azores hydroid fauna and its origin, with discussion of rafting and medusa suppression. Arquipélago. Life and Earth Sciences, 10, 75–99.

    Cornelius, P.F.S. (1995a) North-west European thecate hydroids and their medusae. Part 1. Introduction, Laodiceidae to Haleciidae. Synopses of the British Fauna, n.s., 50, 347 pp.

    Cornelius, P.F.S. (1995b) North-west European thecate hydroids and their medusae. Part 2. Sertulariidae to Campanulariidae. Synopses of the British Fauna, n.s., 50, 386 pp.

    Cornelius, P.F.S. (1998) Taxonomic characters from the hydranths of live thecate hydroids: European Haleciidae (Cnidaria: Leptothecatae). Zoologische Verhandelingen, 323, 79–97.

    Cornelius, P.F.S. & Garfath, J.B. (1980) The coelenterate taxa of Joshua Alder. Bulletin of the British Museum (Natural History), Zoology, 39, 273–291.

    Cornelius, P.F.S. & Ryland, J.S. (1990) Hydrozoa. In: Hayward, P.J. & Ryland, J.S. (Eds.), The marine fauna of the British Isles and north-west Europe. Volume 1. Introduction and protozoans to arthropods. Clarendon Press, Oxford, pp. 107–158.

    Cunha, A.X. da (1944) Hidropólipos das costas de Portugal. Memórias e Estudos do Museu Zoológico da Universidade de Coimbra, 161, 1–101.

    Deshayes, G.P. & Milne Edwards, H. (1836) Histoire naturelle des animaux sans vertèbres, par J.B.P.A. de Lamarck. Deuxième Édition. Tome 2me. J.B. Baillière, Paris, 683 pp.

    Edwards, C. (1965) The hydroid and the medusa Neoturris pileata. Journal of the Marine Biological Association of the United Kingdom, 45, 443–468.

    Edwards, C. (1972) The hydroids and the medusae Podocoryne areolata, P. borealis and P. carnea. Journal of the Marine Biological Association of the United Kingdom, 52, 97–144.

    Edwards, C. (1973a) The medusa Modeeria rotunda and its hydroid Stegopoma fastigiatum, with a review of Stegopoma and Stegolaria. Journal of the Marine Biological Association of the United Kingdom, 53, 573–600.

    Edwards, C. (1973b) The medusa Mitrocomella polydiadema and its hydroid. Journal of the Marine Biological Association of the United Kingdom, 53, 601–607.

    Edwards, C. & Harvey, S.M. (1975) The hydroids Clava multicornis and Clava squamata. Journal of the Marine Biological Association of the United Kingdom, 55, 879–886.

    Ehrenberg, C.G. (1834) Beiträge zur physiologischen Kenntniss der Corallenthiere im allgemeinen, und besonders des rothen Meeres, nebst einem Versuche zur physiologischen Systematik derselben. Abhandlung der Königlichen Akademie der Wissenschaften, Berlin, 1, 225–380.

    Ellis, J. (1755) An essay towards a natural history of the corallines, and other marine productions of the like kind, commonly found on the coasts of Great Britain and Ireland… Printed for the author, London, 104 pp.

    Ellis, J. & Solander, D. (1786) The natural history of many curious and uncommon zoophytes, collected from various parts of the globe by the late John Ellis… Systematically arranged and described by the late Daniel Solander… Benjamin White & Son, and Peter Elmsly, London, 206 pp.

    Eschscholtz, F. (1829) System der Acalephen. Eine ausfürliche Beschreibung aller medusenartigen Strahltiere. Ferdinand Dümmler, Berlin, 190 pp.

    Fenchel, A. (1905) Ueber Tubularia larynx Ellis (T. coronata Abildgaard). Revue Suisse de Zoologie, 13, 507–580.

    Fewkes, J.W. (1882) On the Acalephae of the east coast of New England. I. Medusae from Newport. II. Acalephae collected by the U.S. Fish Commission during the summer of 1880 and 1881. Bulletin of the Museum of Comparative Zoölogy at Harvard College, 9, 291–310.

    Fleming, J. (1820) Observations on the natural history of the Sertularia gelatinosa of Pallas. Edinburgh Philosophical Journal, 2, 82–89.

    Fleming, J. (1823) Gleanings of natural history, gathered on the coast of Scotland during a voyage in 1821. Edinburgh Philosophical Journal, 8, 294–303.

    Fleming, J. (1828) A history of British animals, exhibiting the descriptive characters and systematical arrangement of the genera and species of quadrupeds, birds, reptiles, fishes, Mollusca, and Radiata of the United Kingdom. Bell & Bradfute, Edinburgh, 565 pp.

    Flórez González, L. (1983) Inventario preliminar de la fauna hydroide de la Bahia de Cartagena y areas adyacentes. Boletín del Museo del Mar, Bogota, 11, 112–140.

    Folino-Rorem, N.C., Darling, J.A. & D’Ausilio, C.A. (2009) Genetic analysis reveals multiple cryptic invasive species of the hydrozoan genus Cordylophora. Biological Invasions, 11, 1869–1882.

    Forbes, E. (1846) On the pulmograde medusae of the British seas. Annals and Magazine of Natural History, 18, 284–287.

    Forbes, E. (1848) A monograph of the British naked-eyed medusae: with figures of all the species. Ray Society, London, 104 pp.

    Forbes, E. & Goodsir, J. (1853) On some remarkable marine Invertebrata new to the British seas. Transactions of the Royal Society of Edinburgh, 20, 307–315.

    Forsskål, P. (1775) Descriptiones animalium avium, amphibiorum, piscium, insectorum, vermium; quae in itinere orientali observavit Petrus Forskål. Prof. Haun. Post mortem auctoris edidit Carsten Niebuhr. Mölleri, Hauniae, 140 pp.

    Fraser, C.M. (1912) Some hydroids of Beaufort, North Carolina. Bulletin of the United States Bureau of Fisheries, 30, 339–387.

    Fraser, C.M. (1924) Acaulis primarius Stimpson. Transactions of the Royal Society of Canada, Section V, Series 3, 18, 165–173.

    Fraser, C.M. (1937) Hydroids of the Pacific coast of Canada and the United States. University of Toronto Press, Toronto, 207 pp.

    Fraser, C.M. (1938) Hydroids of the 1934 Allan Hancock Pacific Expedition. Allan Hancock Pacific Expeditions, 4(1), 1–105.

    Fraser, C. M. (1944) Hydroids of the Atlantic coast of North America. University of Toronto Press, Toronto, 451 pp.

    Fraser, C.M. (1948) Hydroids of the Allan Hancock Pacific Expeditions since March, 1938. Allan Hancock Pacific Expeditions, 4(5), 179–343.

    Galea, H.R., Häussermann, V. & Försterra, G. (2009) New additions to the hydroids (Cnidaria: Hydrozoa) from the fjords region of southern Chile. Zootaxa, 2019, 1–28.

    Gislén, T. (1930) Epibioses of the Gullmar Fjord II. Kristinebergs Zoologiska Station 1877–1927, Skriftserie Utgiven av K. Svenska Vetenskapsakademien, 4, 1–380.

    Govindarajan, A.F., Halanych, K.M. & Cunningham, C.W. (2005) Mitochondrial evolution and phylogeography in the hydrozoan Obelia geniculata (Cnidaria). Marine Biology, 146, 213–222.

    Gravier-Bonnet, N. (2007) Polymorphism in hydroids: the extensible polyp of Halecium halecinum (Cnidaria: Hydrozoa: Haleciidae). Journal of the Marine Biological Association of the United Kingdom, 88, 1731–1736.

    Gray, J.E. (1848) List of the specimens of British animals in the collection of the British Museum. Part I. Centroniae or radiated animals. British Museum, London, 173 pp.

    Guenther, J., Carl, C. & Sunde, L.M. (2009) The effects of colour and copper on the settlement of the hydroid Ectopleura larynx on aquaculture nets in Norway. Aquaculture, 292, 252–255.

    Guenther, J., Misimi, E. & Sunde, L.M. (2010) The development of biofouling, particularly the hydroid Ectopleura larynx, on commercial salmon cage nets in mid-Norway. Aquaculture, 300, 120–127.

    Haeckel, E. (1879) Das System der Medusen. Erster Theil einer Monographie der Medusen. Denkschriften der Medicinisch-Naturwissenschaftlichen Gesellschaft zu Jena, 1, 360 pp.

    Hamond, R. (1957) Notes on the Hydrozoa of the Norfolk coast. Journal of the Linnean Society of London, Zoology, 43, 294–324.

    Hancock, D.A., Drinnan, R.E. & Harris, W.N. (1956) Notes on the biology of Sertularia argentea L. Journal of the Marine Biological Association of the United Kingdom, 35, 307–325.

    Hansson, H.G. (1998) NEAT (North East Atlantic Taxa): South Scandinavian marine Cnidaria + Ctenophora Check-List. http://www.tmbl.gu.se/libdb/taxon/neat_pdf/NEAT*Coelenterata.pdf

    Hartlaub, C. (1897) Die Hydromedusen Helgolands. Zweiter Bericht. Wissenschaftliche Meeresuntersuchungen, neue folge, 2, 449–537.

    Hartlaub, C. (1905) Die Hydroiden der magalhaensischen Region und chilenischen Küste. Zoologische Jahrbücher, Supplement-Band 6, Fauna Chilensis, 3, 497–714.

    Hartlaub, C. (1913) XII. Craspedote Medusen. I. Teil. 3. Lief.: Familie IV Tiaridae. Nordisches Plankton, 6, 237–363.

    Hartlaub, C. & Scheuring, L. (1915). Zoologische Ergebnisse einer Untersuchungsfahrt des Deutschen Seefischerei-Vereins nach der Bäreninsel und Westspitzbergen, ausgeführt im Sommer 1898 auf S. M. S. "Olga." Die Hydroiden der Olga-Expedition (1898). Wissenschaftliche Meeresuntersuchungen, Helgoland, neue folge, 11(2), 67–90.

    Hassall, A.H. (1848) Definitions of three new British zoophytes. Zoologist, 6: 2223.

    Heller, C. (1968) Die Zoophyten und Echinodermen des Adriatischen Meeres. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 18, 1–88.

    Henry, L.-A. (2001) Intertidal zonation and seasonality of the marine hydroid Dynamena pumila (Cnidaria: Hydrozoa). Canadian Journal of Zoology, 80, 1526–1536.

    Henry, L.-A. & Kenchington, E. (2004) Differences between epilithic and epizoic hydroid assemblages from commercial scallop grounds in the Bay of Fundy, northwest Atlantic. Marine Ecology Progress Series, 266, 123–134.

    Henry, L.-A., Kenchington, E.L.R., Kenchington, T.J., MacIsaac, K.G., Bourbonnais-Boyce, C. & Gordon, D.C., Jr. (2004) Impacts of otter trawling on colonial epifaunal assemblages on a cobble bottom ecosystem on Western Bank (northwest Atlantic). Marine Ecology Progress Series, 306, 63–78.

    Henry, L.-A., Nizinski, M.S. & Ross, S.W. (2008) Occurrence and biogeography of hydroids (Cnidaria: Hydrozoa) from deep-water coral habitats off the southeastern United States. Deep-Sea Research I, 55, 788–800.

    Hincks, T. (1861) A catalogue of the zoophytes of south Devon and south Cornwall. Annals and Magazine of Natural History, third series, 8, 251–262.

    Hincks, T. (1866) On new British Hydroida. Annals and Magazine of Natural History, third series, 18, 296–299.

    Hincks, T. (1868) A history of the British hydroid zoophytes. John van Voorst, London, 338 pp.

    Hincks, T. (1874a) Notes on Norwegian Hydroida from deep water. Annals and Magazine of Natural History, fourth series, 13, 125–137.

    Hincks, T. (1874b) On deep-water Hydroida from Iceland. Annals and Magazine of Natural History, fourth series, 13, 146–153.

    Hirohito, The Showa Emperor (1995) The hydroids of Sagami Bay. Part II. Thecata. Biological Laboratory, Imperial Household, Tokyo, Japan, 355 pp.

    Hughes, R.G. (1992). Morphological adaptations of the perisarc of the intertidal hydroid Dynamena pumila to reduce damage and enhance feeding efficiency. Scientia Marina, 56, 269–277.

    Hult, J. (1941) Ueber einige für die Westküste Schwedens neue niedere Evertebraten. Zoologiska Bidrag från Uppsala, 20, 455–460.

    International Commission on Zoological Nomenclature (1982) Opinion 1220. Halecium Oken, 1815, (Coelenterata, Hydroida): ruled to be an available name and conserved. Bulletin of Zoological Nomenclature, 45, 155–156.

    International Commission on Zoological Nomenclature (1985) Opinion 1345. Laomedea flexuosa Alder, 1857, Sertularia volubilis Linnaeus, 1758 and Campanularia johnstoni Alder, 1856 designated as type species of Laomedea Lamouroux, 1812, Campanularia Lamarck, 1816 and Clytia Lamouroux, 1812 (Coelenterata, Hydroida) respectively. Bulletin of Zoological Nomenclature, 42, 271–273.

    International Commission on Zoological Nomenclature (1988) Opinion 1485. Filellum serpens Hassall, 1848 (Cnidaria: Hydrozoa): specific and generic names conserved. Bulletin of Zoological Nomenclature, 45, 155–156.

    International Commission on Zoological Nomenclature (2000) Opinion 1956. Eudendrium arbuscula Wright, 1859 (Cnidaria, Hydrozoa): specific name conserved. Bulletin of Zoological Nomenclature, 57, 180–181.

    Jäderholm, E. (1907) Über einige nordische Hydroiden. Zoologischer Anzeiger, 32, 371–376.

    Jäderholm, E. (1909) Northern and Arctic invertebrates in the collection of the Swedish State Museum (Riksmuseum). IV. Hydroiden. Kungliga Svenska Vetenskapsakademiens Handlingar, ny följd, 45(1), 1–124.

    Jäderholm, E. (1919) Zur Kenntnis der Hydroidenfauna Japans. Arkiv för Zoologi, 12(9), 1–34.

    Jägerskiöld, L.A. (1971) A survey of the marine benthonic macro-fauna along the Swedish west coast 1921–1938. Acta Regiae Societatis Scientiarum et Litterarum Gothobergensis, Zoologica, 6, 1–146.

    Johnston, G. (1833) Illustrations in British zoology. 15. Plumularia catharina. Magazine of Natural History, 6, 497–499.

    Johnston, G. (1837) A catalogue of the zoophytes of Berwickshire. History of the Berwickshire Naturalists’ Club, 1, 107–108. [Note: dating of this article follows Cornelius (1995b: 360)].

    Johnston, G. (1838) A history of the British zoophytes. W.H. Lizars, Edinburgh, 341 pp.

    Johnston, G. (1847) A history of the British zoophytes. Second edition. John Van Voorst, London, 488 pp.

    Jonsson, L.G., Nilsson, P.G., Floruta, F. & Lundälv, T. (2004) Distributional patterns of macro- and megafauna associated with a reef of the cold-water coral Lophelia pertusa on the Swedish west coast. Marine Ecology Progress Series, 284, 163–171.

    Kashin, I.A., Bagaveeva, E.V. & Chaplygina, S.F. (2003) Fouling communities of hydrotechnical constructions in Nakhodka Bay (Sea of Japan). Russian Journal of Marine Biology, 29, 267–283.

    Kosevich, I.A. (2006) Changes in the patterning of a hydroid colony. Zoology, 109, 244–259.

    Kramp, P.L. (1911) Report on the hydroids collected by the Danmark Expedition at north-east Greenland. Meddelelser om Grønland, 45, 341–396.

    Kramp, P.L. (1915) Hydroids collected by the “Tjalfe” Expedition to the west coast of Greenland in 1908 and 1909. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn, 66, 1–36.

    Kramp, P.L. (1926) Medusae. Part II. Anthomedusae. Danish Ingolf-Expedition, 5(10), 1–102.

    Kramp, P.L. (1932a) Hydroids collected in west-Greenland fjords in 1911 and 1912. Meddelelser om Grønland, 91(3), 1–35.

    Kramp, P.L. (1932b) The Godthaab Expedition 1928. Hydroids. Meddelelser om Grønland, 79(1), 1–86.

    Kramp, P.L. (1935a) Corydendrium dispar, a new athecate hydroid from Scandinavian seas, with remarks on classification. Göteborgs Kungliga Vetenskaps- och Vitterhets-Samhälles Handlingar, femte följden, Series B, 4(11), 1–15.

    Kramp, P.L. (1935b) Polypdyr (Coelenterata) I. Ferskvandspolypper og goplepolypper. Danmarks Fauna, 41, 1–207.

    Kramp, P.L. (1938) Marine Hydrozoa. a. Hydroida. Zoology of Iceland, 2 (Part 5a), 1–82.

    Kramp, P.L. (1941) Notes on the hydroid Campanulina panicula G. O. Sars. Göteborgs Kungliga Vetenskaps- och Vitterhets-Samhälles Handlingar, sjätte följden, Series B, 1(2), 1–11.

    Kramp, P.L. (1942) V. Marine Hydrozoa. Zoology of the Faroes, 1(1), 1–59.

    Kramp, P.L. (1943) The zoology of east Greenland. Hydroida. Meddelelser om Grønland, 121(11), 1–52.

    Kramp, P.L. (1947) Medusae Part III. Trachylina and Scyphozoa with zoogeographical remarks on all the medusae of the northern Atlantic. Danish Ingolf-Expedition, 5(14), 1–66.

    Kramp, P.L. (1961) Synopsis of the medusae of the world. Journal of the Marine Biological Association of the United Kingdom, 40, 1–469.

    Kudelin, N.V. (1914) Gidroidy (Hydroidea). Tom II. Plumulariidae, Campanulinidae i Sertulariidae. Fauna Rossii i Supredelnykh Stran, 2(2), 139–526.

    Kühn, A. (1913) Entwicklungsgeschichte und Verwandtschaftsbeziehungen der Hydrozoen. I. Teil: Die Hydroiden. Ergebnisse und Fortschritte der Zoologie, 4, 1–284.

    Lamouroux, J.V.F. (1812) Extrait d’un mémoire sur la classification des polypiers coralligènes non entièrement pierreux. Nouveau Bulletin des Sciences, par la Société Philomatique de Paris, 3, 181–188.

    Lamouroux, J.V.F. (1816) Histoire des polypiers coralligènes flexibles, vulgairement nommés zoophytes. F. Poisson, Caen, 560 pp.

    Lamouroux, J.V.F. (1821) Exposition méthodique des genres de l’ordre des polypiers, avec leur description et celle des principales espèces, figurées dans 84 planches; les 63 premières appartenant à l’histoire naturelle des zoophytes d’Ellis et Solander. Agasse, Paris, 115 pp.

    Lamouroux, J.V.F. (1824) Description des polypiers flexibles. In: Quoy, J.R.C. & Gaimard, J.P. (Eds.), Zoologie. Voyage autour du monde, entrepris par ordre du Roi, exécuté sur les corvettes de S.M. l'Uranie et la Physicienne, pendant les années 1817, 1818, 1819 et 1820; par M. Louis de Freycinet. Pillet Aîné, Paris, 603–643.

    Latham, E. (1963) The hydroid Leuckartiara octona (Fleming) and its association with the polychaete Aphrodite aculeata (L.). Annals and Magazine of Natural History, series 13, 5, 523–528.

    Leclère, L., Schuchert, P. & Manuel, M. (2007) Phylogeny of the Plumularioidea (Hydrozoa, Leptothecata): evolution of colonial organization and life cycle. Zoologica Scripta, 36, 371–394.

    Leloup, E. (1934) Contributions à l’étude de la faune Belge. V. – Les hydropolypes épizoïques du ver polychète, Aphrodite aculeata (Linné). Bulletin du Musée Royal d’Histoire Naturelle de Belgique, 10 (41), 1–6.

    Leloup, E. (1974) Hydropolypes calyptoblastiques du Chili. Report No. 48 of the Lund University Chile Expedition 1948–1949. Sarsia, 55, 1–61.

    Lendenfeld, R. von (1885) The Australian hydromedusae. Part I. The classification of the Hydromedusae. Proceedings of the Linnean Society of New South Wales, 9, 206–241.

    Lesson, R.P. (1830) Voyage autour du monde, pendant les années 1822, 1823, 1824 et 1825. Zoologie. Description des zoophytes échinodermes. Voyage autour du monde, éxecuté par ordre du Roi, sur la corvette de sa majesté, La Coquille, pendant les années 1822, 1823, 1824 et 1825. Histoire Naturelle, Zoologie, Tome 2, Partie 2, Chapitre 14. Arthus Bertrand, Paris, 155 pp.

    Levinsen, G.M.R. (1893a) Om fornyelsen af ernæringsindividerne hos hydroiderne. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn, series 5, 4, 14–31.

    Levinsen, G.M.R. (1893b) Meduser, ctenophorer og hydroider fra Grønlands vestkyst, tilligemed bemærkninger om hydroidernes systematik. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn, series 5, 4, 143–220.

    Levinsen, G.M.R. (1893c) Om en ny Thujaria-Art fra Kara-Havet. Thujaria carica, nov. sp. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn, series 5, 4, 213–214.

    Lindström, G. (1856a) Bidrag till kännedomen om Östersjöns invertebrat-fauna. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 12, 49–73.

    Lindström, G. (1856b) Om utvecklingen af Sertularia pumila L. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 12, 365–374.

    Linko, A.K. (1911) Gidroidy (Hydroidea). Tom I. Haleciidae, Lafoëidae, Bonneviellidae i Campanulariidae. Fauna Rossii i Supredelnykh Stran, 1, 1–250.

    Linko, A.K. (1912) Gidroidy (Hydroidea). Tom II. Plumulariidae, Campanulinidae i Sertulariidae. Fauna Rossii i Supredelnykh Stran, 2(1), 1–138.

    Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Laurentii Salvii, Holmiae, 823 pp.

    Linnaeus, C. (1761) Fauna Svecica sistens animalia Sveciae regni: Mammalia, Aves, Amphibia, Pisces, Insecta, Vermes. Distributa per classes & ordines, genera & species, cum differentiis specierum, synonymis auctorum, nominibus incolarum, locis natalium, descriptionibus insectorum. Laurentii Salvii, Stockholmiae, 578 pp.

    Linnaeus, C. (1767) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Pars II. Editio duodecima, reformata. Laurentii Salvii, Holmiae, pp. 533–1317.

    Lönnberg, E. (1898) Undersökningar rörande Öresunds djurlif. Meddelanden från Kongl. Landtbruksstyrelsen, år 1898, 43, 1–77.

    Lönnberg, E. (1899a) Clava glomerata mihi, eine anscheinend neue Hydroide. Zoologischer Anzeiger 22: 45–46.

    Lönnberg, E. (1899b) Fortsatta undersökningar rörande Öresunds djurlif. Meddelanden från Kongl. Landtbruksstyrelsen, år 1899, 49, 1–24.

    Lönnberg, E. (1902) Några smärre iakttagelser rörande faunan i Bohuslän i mars månad 1902. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 59, 169–174.

    Lönnberg, E. (1903) Undersökningar rörande Skeldervikens och Angränsande Kattegat-områdes djurlif på Förordnande af Kongl. Landtbruksstyrelsen. Meddelanden från Kongl. Landtbruksstyrelsen, år 1902, 80, 1–70.

    Lorenz, L. von (1886) Polypomedusen von Jan Mayen. Gesammelt von Dr. F. Fischer. Die Internationale Polarforschung 1882–1883. Die Österreichische Polarstation Jan Mayen, 3, 25–28.

    Lovén, S. (1836) Bidrag till kännedomen af slägtena Campanularia och Syncoryna. Kongl. Vetenskaps-Academiens Handlingar, för År 1835, 260–281.

    Lovén, S. (1858) Till utvecklingen af Hydractinia. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 14, 305–313.

    Lundin, K., Andréasson, E., Högström, C.A. & Karlsson, A. (2007) Faunistiskt nytt 2006 – marina evertebrater. Göteborgs Naturhistoriska Museum Årstryck, 2007, 31–45.

    Lundin, K., Karlsson, A., Möller, P., Högström, C.A. & Andréasson, E. (2009) Faunistiskt nytt 2008 – marina evertebrater. Göteborgs Naturhistoriska Museum Årstryck, 2009, 31–46.

    Lütken, C. (1850) Nogle Bemaerkninger om Medusernes systematiske Inddeling, navnlig med Hensyn til Forbes’s History of brittish naked-eyed medusae. Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjöbenhavn, 1850, 15–35.

    Marfenin, N.N. (1983) Novyĭ vid Cordylophora (Hydrozoa, Clavidae) iz Chernogo Morya. A new species of the genus Cordylophora (Hydrozoa, Clavidae) from the Black Sea. Zoologicheskii Zhurnal, 62, 1732–1734.

    Marfenin, N.N. & Belorustseva, S.A. (2008) How the distribution of colonies of the hydroid Laomedea flexuosa is limited to a narrow belt along the lower littoral zone. Journal of the Marine Biological Association of the United Kingdom, 88, 1559–1566.

    Marktanner-Turneretscher, G. (1890) Die Hydroiden des K. K. Naturhistorischen Hofmuseums. Annalen des K. K. Naturhistorischen Hofmuseums, 5, 195–286.

    Marktanner-Turneretscher, G. (1895) Zoologische Ergebnisse der im Jahre 1889 auf Kosten der Bremer geographischen Gesellschaft von Dr. Willy Kükenthal und Dr. Alfred Walter ausgeführten Expedition nach Ost-Spitzbergen. Hydroiden. Zoologische Jahrbücher, Abtheilung für Systematik, Geographie und Biologie der Thiere, 8, 391–438.

    Marques, A.C., Mergner, H., Höinghaus, R., Santos, C.M.D., & Vervoort, W. (2000) Morphological study and taxonomical notes on Eudendriidae (Cnidaria: Hydrozoa: Athecatae/Anthomedusae). Zoologische Mededelingen, 74, 75–118.

    Marques, A.C., Peña Cantero, A.L., Miranda, T.P. & Migotto, A.E. (2011) Revision of the genus Filellum Hincks, 1868 (Lafoeidae, Leptothecata, Hydrozoa). Zootaxa, 3129, 1–28.

    Marques A.C., & Vervoort, W. (1999) Eudendrium arbuscula Wright, 1859 (Cnidaria, Hydrozoa): proposed conservation of the specific name. Bulletin of Zoological Nomenclature, 56, 16–18.

    McCrady, J. (1859) Gymnopthalmata of Charleston Harbor. Proceedings of the Elliott Society of Natural History, 1, 103–221.

    Medel, M.D. & Vervoort, W. (1995) Plumularian hydroids (Cnidaria: Hydrozoa) from the Strait of Gibraltar and nearby areas. Zoologische Verhandelingen, 300, 1–72.

    Medel, M.D., García, F.G. & Vervoort, W. (1998) The family Haleciidae (Cnidaria: Hydrozoa) from the Strait of Gibraltar and nearby areas. Zoologische Mededelingen, 72, 29–50.

    Medel, M.D. & Vervoort, W. (1998) Atlantic Thyroscyphidae and Sertulariidae (Hydrozoa, Cnidaria) collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, The Netherlands. Zoologische Verhandelingen, 320, 1–85.

    Medel, M.D. & Vervoort, W. (2000) Atlantic Haleciidae and Campanulariidae (Hydrozoa, Cnidaria) collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, The Netherlands. Zoologische Verhandelingen, 330, 1–68.

    Michanek, G. (1967) Quantitative sampling of benthic organisms by diving on the Swedish west coast. Helgoländer Wissenschaftliche Meeresuntersuchungen, 15, 455–459.

    Migotto, A.E. & Andrade, L.P. de (2000) The life cycle of Hebella furax (Cnidaria: Hydrozoa): a link between a lafoeid hydroid and a laodiceid medusa. Journal of Natural History, 34, 1871–1888.

    Millard, N.A.H. (1959) Hydrozoa from ship’s hulls and experimental plates in Cape Town docks. Annals of the South African Museum, 45, 239–256.

    Millard, N.A.H. (1962) The Hydrozoa of the south and west coasts of South Africa. Part I. The Plumulariidae. Annals of the South African Museum, 46, 261–319.

    Millard, N.A.H. (1975) Monograph on the Hydroida of southern Africa. Annals of the South African Museum, 68, 1–513.

    Mortensen, P.B. & Fosså, J.H. (2006) Species diversity and spatial distribution of invertebrates on deep-water Lophelia reefs in Norway. Proceedings of the 10th International Coral Reef Symposium, 1849–1868.

    Moura, C.J., Harris, D.J., Cunha, M.R. & Rogers, A.D. (2008) DNA barcoding reveals cyptic diversity in marine hydroids (Cnidaria, Hydrozoa) from coastal and deep-sea environments. Zoologica Scripta, 37, 93–108.

    Moura, C.J., Cunha, M.R., Porteiro, F.M. & Rogers, A.D. (2011) The use of the DNA barcode gene 16S mRNA for the clarification of taxonomic problems within the family Sertulariidae (Cnidaria, Hydrozoa). Zoologica Scripta, 40, 520–537.

    Müller, O.F. (1776) Zoologiae Danicae prodromus, seu animalium Daniae et Norvegiae indigenarum characteres, nomina, et synonyma imprimis popularium. Hallageriis, Hauniae, 282 pp.

    Naumov, D.V. (1960) Gidroidy i gidromeduzy morskikh, solonovatovodnykh i presnovodnykh basseinov SSSR. Akademiya Nauk SSSR, Opredeliteli po Faune SSSR, 70, 1–626.

    Norman, A.M. (1864) On undescribed British Hydrozoa, Actinozoa, and Polyzoa. Annals and Magazine of Natural History, series 3, 13, 82–90.

    Norman, A.M. (1867) Report of the committee appointed for the purpose of exploring the coasts of the Hebrides by means of the dredge.–Part II. On the Crustacea, Echinodermata, Polyzoa, Actinozoa, and Hydrozoa. Report of the British Association for the Advancement of Science, Nottingham, 1866, 193–206.

    Nutting, C.C. (1896) Notes on Plymouth hydroids. Journal of the Marine Biological Association of the United Kingdom, new series, 4, 146–154.

    Nutting, C.C. (1899) Hydroida from Alaska and Puget Sound. Proceedings of the United States National Museum, 21, 741–753.

    Nutting, C.C. (1900) American hydroids. Part I. The Plumularidae. Smithsonian Institution, United States National Museum Special Bulletin, 4(1), 1–285.

    Nutting, C.C. (1904) American hydroids. Part II. The Sertularidae. Smithsonian Institution, United States National Museum Special Bulletin, 4(2), 1–325.

    Oken, L. (1815) Okens Lehrbuch der Naturgeschichte. III. Theil. Zoologie. Volume 1. Oken, Jena, 842 pp.

    Orbigny, A. d’ (1846) Voyage dans l’Amérique Méridionale (le Brésil, la République orientale de l’Uruguay, la République Argentine, La Patagonie, la République du Chili, la République de Bolivia, La République du Pérou), exécuté pendant les années 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833. Tome 5, 4e Partie: zoophytes. Levrault, Strasbourg, 28 pp.

    Osburn, H.L. (1893) Biological descriptions of certain common hydroid animals. American Monthly Microscopical Journal, 14, 94–100.

    Östman, C. (1979a) Two types of nematocysts in Campanulariidae (Cnidaria, Hydrozoa) studied by light and scanning electron microscopy. Zoologica Scripta, 8, 5–12.

    Östman, C. (1979b) Nematocysts in the Phialidium medusae of Clytia hemisphaerica (Hydrozoa, Campanulariidae) studied by light and scanning electron microscopy. Zoon, 7, 125–142.

    Östman, C. (1982a) Isoenzymes and taxonomy in Scandinavian hydroids (Cnidaria, Campanulariidae). Zoologica Scripta, 11, 155–163.

    Östman, C. (1982b) Nematocysts and taxonomy in Laomedea, Gonothyraea and Obelia (Hydrozoa, Campanulariidae). Zoologica Scripta, 11, 227–241.

    Östman, C. (1983) Taxonomy of Scandinavan hydroids (Cnidaria, Campanulariidae): a study based on nematocyst morphology and isoenzymes. Acta Universitatis Upsaliensis, 672, 1–22.

    Östman, C. (1987) New techniques and old problems in hydrozoan systematics. In: Bouillon, J., Boero, F., Cicogna, F. & Cornelius, P.F.S. (Eds.), Modern trends in the systematics, ecology, and evolution of hydroids and hydromedusae. Clarenden Press, Oxford, pp. 67–82.

    Östman, C. (1999) Nematocysts and their value as taxonomic parameters within the Campanulariidae (Hydrozoa). A review based on light and scanning electron microscopy. Zoosystematica Rossica, Supplement 1, 17–28.

    Östman, C., Myrdal, M., Nyvall, P., Lindström, J., Björklund, M. & Aguirre, A. (1995) Nematocysts in Tubularia larynx (Cnidaria, Hydrozoa) from Scandinavia and the northern coast of Spain. Scientia Marina, 59, 165–179.

    Östman, C., Piraino, S. & Roca, I. (1987) Nematocyst comparisons between some Mediterranean and Scandinavian campanulariids (Cnidaria, Hydrozoa). In: Bouillon, J., Boero, F., Cicogna, F. & Cornelius, P.F.S. (Eds.), Modern trends in the systematics, ecology, and evolution of hydroids and hydromedusae. Clarenden Press, Oxford, pp. 299–310.

    Owen, R. (1843) Lectures on the comparative anatomy and physiology of the invertebrate animals: delivered at the Royal College of Surgeons, in 1843. Longman, Brown, Green, & Longmans, London, 392 pp.

    Pallas, P.S. (1866) Elenchus zoophytorum sistens generum adumbrationes generaliores et specierum cognitarum succinctas descriptiones cum selectis auctorum synonymis. Hagae, Franciscum Varrentrapp, 451 pp.

    Pallas, P.S. (1771) Reise durch verschiedene Provinzen des russischen Reichs. Erster Theil. Kayserliche Academie der Wissenschaften, St. Petersburg, 504 pp.

    Peña Cantero, Á.L. & García Carrascosa, A.M. (1999) Biogeographical distribution of the benthic thecate hydroids collected during the Spanish “Antártida 8611” Expedition and comparison between Antarctic and Magellan benthic hydroid faunas. Scientia Marina, 63 (supl. 1), 209–218.

    Peña Cantero, Á.L., García Carrascosa, A.M. & Vervoort, W. (1998) On the species of Filellum Hincks, 1868 (Cnidaria: Hydrozoa) with the description of a new species. Journal of Natural History, 32, 297–315.

    Peña Cantero, Á.L., Sentandreu, V. & Latorre, A. (2010) Phylogenetic relationships of the endemic Antarctic benthic hydroids (Cnidaria, Hydrozoa): what does the mitochondrial 16S rRNA tell us about it? Polar Biology, 33, 41–57.

    Peña Cantero, Á.L. & Vervoort, W. (1999) Review of the genus Schizotricha Allman, 1883 (Cnidaria, Hydrozoa, Halopterididae). Journal of Natural History, 33, 351–386.

    Péron, F. & Lesueur, C.A. (1810) Tableau des caractères génériques et spécifiques de toutes les espèces de méduses connues jusqu’à ce jour. Annales du Muséum d’Histoire Naturelle, 14, 325–366.

    Petersen, K.W. (1979) Development of coloniality in Hydrozoa. In: Larwood, G. & Rosen, B.R. (Eds.), Biology and systematics of colonial animals. Academic Press, New York, pp. 105–139.

    Petersen, K.W. (1990) Evolution and taxonomy in capitate hydroids and medusae (Cnidaria: Hydrozoa). Zoological Journal of the Linnean Society, 100, 101–231.

    Picard, J. (1952) Note sur les hydraires littoraux de Banyuls-sur-Mer. Vie et Milieu, 2, 338–349.

    Pieper, F.W. (1884) Ergänzungen zu “Heller’s Zoophyten etc. des Adriatischen Meeres.” Zoologischer Anzeiger, 7, 164–169.

    Quoy, J.R.C. & Gaimard, J.P. (1827) Observations zoologiques faites à bord de l’Astrolabe, en mai 1826, dans le Détroit de Gibraltar. Annales des Sciences Naturelles, 10, 172–193.

    Ramil, F., & Iglesias, A. (1988) Sobre la presencia de Opercularella panicula (Sars, 1873) (Cnidaria: Hydroida) en las costas de la Peninsula Iberica. Thalassas, 6, 79–82.

    Ramil, F., Parapar, J. & Vervoort, W. (1992) The genus Sertularella Gray, 1848 (Cnidaria: Hydroida) along the coasts of Galicia (Spain). Zoologische Mededelingen, 66, 493–524.

    Ramil, F. & Vervoort, W. (1992) Report on the Hydroida collected by the “BALGIM” expedition in and around the Strait of Gibraltar. Zoologische Verhandelingen, 277, 1–262.

    Ramil, F., Vervoort, W. & Ansín, J.A. (1998) Report on the Haleciidae and Plumularioidea (Cnidaria, Hydrozoa) collected by the French SEAMOUNT 1 expedition. Zoologische Verhandelingen, 322, 1–42.

    Rees, W.J. (1938) Observations on British and Norwegian hydroids and their medusae. Journal of the Marine Biological Asociation of the United Kingdom, 23, 1–42.

    Rees, W.J. (1941) The hydroid of the medusa Cosmetira pilosella Forbes. Proceedings of the Royal Society of Edinburgh, Section B (Biology), 61, 55–58.

    Rees, W.J. (1956a) A revision of the hydroid genus Perigonimus M. Sars, 1846. Bulletin of the British Museum (Natural History), Zoology, 3, 337–350.

    Rees, W.J. (1956b) A revision of some northern gymnoblastic hydroids in the Zoological Museum, Oslo. Nytt Magasin for Zoologi, 4, 109–120.

    Rees, W.J. & Rowe, M. (1969) Hydroids of the Swedish west coast. Acta Regiae Societatis Scientiarum et Litterarum Gothobergensis, Zoologica, 3: 1–23.

    Rees, W.J. & Vervoort, W. (1987) Hydroids from the John Murray Expedition to the Indian Ocean, with revisory notes on Hydrodendron, Abietinella, Cryptolaria and Zygophylax (Cnidaria: Hydrozoa). Zoologische Verhandelingen, 237, 1–209.

    Reyes, R. & Campos, N.H. (1992) Macroinvertebrados colonizadores de raíces de Rhizophora mangle en la Bahía de Chengue, Caribe Colombiano. Boletín de Investigaciones Marinas y Costeras–INVEMAR, 21, 101–116.

    Ritchie, J. (1911) Contribution to our knowledge of the hydroid fauna of the west of Scotland. Being an account of collections made by Sir John Murray, K.C.B., on S.Y. “Medusa.’ Annals of Scottish Natural History, 20(80), 217–225.

    Romanes, G.J. (1876) An account of some new species, varieties, and monstrous forms of medusae. Journal of the Linnean Society, Zoology, 12, 524–531.

    Ronowicz, M. (2007) Benthic hydroids (Cnidaria: Hydrozoa) from Svalbard waters—biodiversity and distribution. Journal of the Marine Biological Association of the United Kingdom, 87, 1089–1094.

    Ronowicz, M., Wlodarska-Kowalczuk, M. & Kuklinski, P. (2008) Factors influencing hydroids (Cnidaria: Hydrozoa) biodiversity and distribution in Arctic kelp forest. Journal of the Marine Biological Association of the United Kingdom, 88, 1567–1575.

    Rossi, S., Gili, J.M. & Hughes, R.G. (2000) The effects of exposure to wave action on the distribution and morphology of the epiphytic hydrozoans Clava multicornis and Dynamena pumila. Scientia Marina, 64 (supl. 1), 135–140.

    Russell, F.S. (1940) On the nematocysts of hydromedusae. III. Journal of the Marine Biological Association of the United Kingdom, 24, 515–523.

    Russell, F.S. (1953) The medusae of the British Isles. Anthomedusae, Leptomedusae, Limnomedusae, Trachymedusae and Narcomedusae. Cambridge University Press, Cambridge, 530 pp.

    Sæmundsson, B. (1902) Bidrag til kundskaben om de islandske hydroider. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn, series 6, 4, 143–220.

    Sars, G.O. (1874) Bidrag til kundskaben om norges hydroider. Forhandlinger i Videnskabs-Selskabet i Christiania, 1873, 91–150.

    Sars, M. (1846) Fauna littoralis Norvegiae oder Beschreibung und Abbildungen neuer oder wenig bekannten Seethiere, nebst Beobachtungen über die Organisation, Lebensweise und Entwickelung derselben. Heft I. Johann Dahl, Christiania, 94 pp.

    Sars, M. (1850) Beretning om en i Sommeren 1849 foretagen zoologisk Reise i Lofoten og Finmarken. Nyt Magazin for Naturvidenskaberne, 6, 121–211. [Note: dating of this article follows Sherborn (1926: 2527), with additional details in Cornelius (1977: 524, 1982: 137)].

    Sars, M. (1857) Bidrag til kundskaben om Middelhavets littoral-fauna, reisebemærkninger fra Italien. Nyt Magazin for Naturvidenskaberne, 9, 110–164.

    Sars, M. (1863) Bemærkninger over fire norske hydroider. Forhandlinger i Videnskabs-Selskabet i Christiania, Aar 1862, 25–39.

    Schönborn, C., Arndt, E.A. & Gosselck, F. (1993) Bestimmungsschlüssel der benthischen Hydrozoen der Ostsee. Mitteilungen aus dem Zoologischen Museum in Berlin, 69, 201–253.

    Schuchert, P. (1997) Review of the family Halopterididae (Hydrozoa, Cnidaria). Zoologische Verhandelingen, 309, 1–162.

    Schuchert, P. (2000) Hydrozoa (Cnidaria) of Iceland collected by the BIOICE programme. Sarsia, 85, 411–438.

    Schuchert, P. (2001a) Hydroids of Greenland and Iceland (Cnidaria, Hydrozoa). Meddelelser om Grønland, Bioscience, 53, 1–184.

    Schuchert, P. (2001b) Survey of the family Corynidae (Cnidaria, Hydrozoa). Revue Suisse de Zoologie, 108, 739–878.

    Schuchert, P. (2003) Hydroids (Cnidaria, Hydrozoa) of the Danish expedition to the Kei Islands. Steenstrupia, 27, 137–256.

    Schuchert, P. (2004) Revision of the European athecate hydroids and their medusae (Hydrozoa, Cnidaria): families Oceanidae and Pachycordylidae. Revue Suisse de Zoologie, 111, 315–369.

    Schuchert, P. (2005) Taxonomic revision and systematic notes on some Halecium species (Cnidaria, Hydrozoa). Journal of Natural History, 39, 607–639.

    Schuchert, P. (2006) The European athecate hydroids and their medusae (Hydrozoa, Cnidaria): Capitata Part 1. Revue Suisse de Zoologie, 113, 325–410.

    Schuchert, P. (2007) The European athecate hydroids and their medusae (Hydrozoa, Cnidaria): Filifera Part 2. Revue Suisse de Zoologie, 114, 195–396.

    Schuchert, P. (2008a) The European athecate hydroids and their medusae (Hydrozoa, Cnidaria): Filifera Part 3. Revue Suisse de Zoologie, 115, 221–302.

    Schuchert, P. (2008b) The European athecate hydroids and their medusae (Hydrozoa, Cnidaria): Filifera Part 4. Revue Suisse de Zoologie, 115, 677–757.

    Schuchert, P. (2010) The European athecate hydroids and their medusae (Hydrozoa, Cnidaria): Capitata Part 2. Revue Suisse de Zoologie, 117, 337–555.

    Schuchert, P. (2011) World Hydrozoa database. Available from: http://www.marinespecies.org/hydrozoa [last consulted on 18 February 2011].

    Schydlowsky, A. (1902) Matériaux relatifs à la faune des polypes hydraires des mers arctiques. I—Les hydraires de la Mer Blanche le long du littoral des Jles Solowetsky. Travaux de la Société des Naturalistes à l’Université Impériale de Kharkov, 36, 1–276.

    Segerstedt, M. (1889) Bidrag till kännedomen om hydroid-faunan vid Sveriges vestkust. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar, 14, Afdelning 4, 4, 1–28.

    Sherborn, C.D. (1926) Index animalium. Section 2, Part 10. Index Eurystomusfunereus. British Museum (Natural History), London, 2249–2568.

    Shih, C.-T., Figueira, A.J.G. & Grainger, E.H. (1971) A synopsis of Canadian marine zooplankton. Bulletin of the Fisheries Research Board of Canada, 176, 1–264.

    Stebbing, A.R.D. (1971) The epizoic fauna of Flustra foliacea [Bryozoa]. Journal of the Marine Biological Association of the United Kingdom, 51, 283–300.

    Stechow, E. (1913) Hydroidpolypen der japanischen Ostküste. II. Teil: Campanularidae, Halecidae, Lafoeidae, Campanulinidae und Sertularidae, nebst Ergänzungen zu den Athecata und Plumularidae. Abhandlungen der Mathematisch-Physikalischen Klasse der Königlichen Bayerischen Akademie der Wissenschaften, 3, Supplement-Band 2, 1–162.

    Stechow, E. (1919a) Zur Kenntnis der Hydroidenfauna des Mittelmeeres, Amerikas und anderer Gebiete, nebst Angaben über einige Kirchenpauer’sche Typen von Plumulariden. Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere, 42, 1–172.

    Stechow, E. (1919b) Neue Ergebnisse auf dem Gebiete der Hydroidenforschung. Münchener Medizinische Wochenschrift, 66, 852–853.

    Stechow, E. (1921a) Neue Genera und Species von Hydrozoen und anderen Evertebraten. Archiv für Naturgeschichte, Abteilung A. 3. Heft, 87, 248–265.

    Stechow, E. (1921b) Über Hydroiden der Deutschen Tiefsee-Expedition, nebst Bemerkungen über einige andre Formen. Zoologischer Anzeiger, 53, 223–236.

    Stechow, E. (1923) Zur Kenntnis der Hydroidenfauna des Mittelmeeres, Amerikas und anderer Gebiete. II. Teil. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 47, 29–270.

    Stechow, E. (1925) Hydroiden der Deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Da