Abstract
Hyptia deansi sp. nov. represents the first record of Evaniidae (Hymenoptera) from Lower Miocene to Upper Oligocene Mexi-can amber, Simojovel, Chiapas, Mexico and is described based on a well preserved female specimen. Phylogenetically relevantmorphological characters are discussed with reference to fossil and extant genera of Evaniidae. The new fossil is placed in theextant New World genus Hyptia Illiger 1807, based on the presence of just a single fore wing cell, the absence of tubular veins M+CU, 1CUa, 1Cub, and the presence of 11 flagellomeres.References
Alonso, J., Arillo, A., Barrón, E., Carmelo Corral, J., Grimalt, J., López, J.F., López, R., Martíne-Delclòs, X., Ortuño, V., Peñalver, E. & Trincão, P.R. (2000) A new fossil resin with biological inclusions in Lower Cretaceous deposits from Alava (northern Spain, Basque-Cantabrian Basin). Journal of Paleontology, 74, 158–178.
Basibuyuk, H.H., Fitton, M.G., Rasnitsyn, A.P. & Quicke, D.L.J. (2000a) Two new genera of the Evaniidae (Insecta: Hymenoptera) from Late Cretaceous New Jersey amber. In: Grimaldi D. (ed.). Studies on Fossils in Amber, with Particular Reference to the Cretaceous of New Jersey, 313–325. Backhuys Publishers; Leiden, The Netherlands; viii + 498 pp.
Basibuyuk, H.H., Fitton, M.G., Rasnitsyn, A.P. & Quicke, D.L.J. (2000b) An archaic new genus of Evaniidae (Insecta: Hymenoptera) and implications for the biology of ancestral evanioids. Bulletin of the Natural History Museum, London (Geology), 56(1), 53–58.
Basibuyuk, H.H., Fitton, M.G., Rasnitsyn, A.P. & Quicke, D.L.J. (2002) The limits of the family Evaniidae (Insecta: Hymenoptera) and a new genus from Lebanese amber. Insect Systematics and Evolution, 33(1), 23–34.
Berggren, W.A. & Van Couvering, J.A. (1974) The late Neogene. Palaeogeography Palaeoclimatology and Palaeoecology, 16, 1–216.
Brues, C.T. (1933) The parasitic Hymenoptera of the Baltic amber, Part 1. Bernstein Forschungen, 3, 4–178.
Deans, A.R. (2005) Annotated catalog of the world’s ensign wasp species (Hymenoptera: Evaniidae). Contributions of the American Entomological Institute, 34, 1–164.
Deans, A.R., Huben, M. (2003) Annotated key to the ensign wasp (Hymenoptera: Evaniidae) genera of the world, with descriptions of three new genera. Proceedings of the Entomological Society of Washington, 105, 859–875.
Deans, A.R., Basibuyuk, H.H., Azar, D. & Nel A. (2004) Descriptions of two new Early Cretaceous (Hauterivian) ensign wasp genera (Hymenoptera: Evaniidae) from Lebanese amber. Cretaceous Research, 25, 509–516.
Deans, A.R., Yoder, M.J. & Dole, K. (2012) Evanioidea Online - catalog of information about evanioid wasps (Hymenoptera). [http://evanioidea.info] (Accessed 16 Jan 2012).
Engel, M.S. (2006) Two ensign wasps in Cretaceous amber from New Jersey and Myanmar (Hymenoptera: Evaniidae). Polskie Pismo Entomologiczne (Polish Journal of Entomology), 75, 443–454.
Hymenoptera Anatomy Consortium (2012) Available at http://glossary.hymao.org. (Accessed on 20 Feb 2012.)
Illiger, J.C.W. 1807. Fauna Etrusca sistens Insecta quae in provinciis Florentina et Pisana praesertim collegit Petrus Rossius, Tomus Secundus. C. G. Fleckeisen, Helmstadii. 511 pp. [in Latin]
Latreille, P.A. (1802) Histoire Naturelle, Générale et Particulière des Crustacés et des Insectes. Ouvrage faisant suite à l’histoire naturelle générale et particulière, composée par Leclerc de Buffon, et rédigée par C.S. Sonnini, membre de plusiers sociétés savantes. Tome troisième. Dufart; Paris, France; xii + 467 pp.
Nel, A., Martíne-Delclòs, X. & Azar, D. (2002a) A new ensign-fly from the Lower-Middle Miocene Dominican amber (Hymenoptera, Evaniidae). Bulletin de la Société Entomologique de France, 107(3), 217–221.
Nel, A., Waller, A., Hodebert, G. & De Ploëg, G. (2002b) An ensign-fly in the lowermost Eocene amber from the Paris basin (Hymenoptera: Evaniidae). Annales de la Société Entomologique de France (N.S.), 38, 263–269.
Peñalver, E., Ortega-Blanco, J., Nel, A. & Delclòs, X. (2010) Mesozoic Evaniidae (Insecta: Hymenoptera) in Spanish Amber: Reanalysis of the Phylogeny of the Evanioidea. Acta Geologica Sinica—English Edition, 84, 809–827.
Poinar, G. & Brown, A.E. (2002) Hymenaea mexicana sp. nov. (Leguminosae: Caesalpinioideae) from Mexican amber indicates Old World connections. Botanical journal of the Linnean Society, 139, 125–132.
Rasnitsyn, A.P. (1975) Hymenoptera Apocrita of Mesozoic. Trudy Paleontologicheskogo Instituta Academii Nauk SSSR (Academy of Sciences of the USSR Transactions of the Palaeontological Institute), 147, 1–134 + plates I-VIII. [in Russian]
Rasnitsyn, A.P. (1991) Early Cretaceous evaniomorphous hymenopteran familes Stigmaphronidae and Cretevaniidae and the subfamily Kotujellitinae (Gasteruptiidae). Paleontologicheskiy Zhurnal, 4, 128–132. [In Russian; English translation in: Paleontological Journal, 25, 172–179]
Rasnitsyn, A.P. & Brothers, D.J. (2007) Two new hymenopteran fossils from the mid-Cretaceous of southern Africa (Hymenoptera: Jurapriidae, Evaniidae). African Invertebrates, 48, 193–202.
Rasnitsyn A.P., Jarzembowski, E.A. & Ross, A.J. (1998) Wasps (Insecta: Vespida = Hymenoptera) from the Purbeck and Wealden (Lower Cretaceous) of southern England and their biostratigraphical and palaeoenvironmental signifcance. Cretaceous Research, 19, 329–319.
Sawoniewicz, J. & Kupryjanowicz, J. (2003) Evaniella eocenica sp. nov. from the Baltic amber (Hymenoptera: Evaniidae). Acta Zoologica Cracoviensia ,46 (suppl.), 267–270.
Solórzano Kraemer, M.M. (2007) Systematic, palaeoecology, and palaeobiogeography of the insect fauna from Mexican amber. Palaeontographica Abt. A 282, 1–133.
Zhang, H. & Zhang, J.-F. (2000) A new genus and two new species of Hymenoptera (Insecta) from the Upper Jurassic Yixian formation of Beipiao, Western Liaoning. Acta Micropalaeontologica Sinica, 17, 286–290 + plate I.
Zhang, H., Rasnitsyn, A.P., Wang, D. & Zhang, Y. (2007) Some hatchet wasps (Hymenoptera, Evaniidae) from the Yixian Formation of western Liaoning, China. Cretaceous Research, 28, 310–316.