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Published: 2010-08-31
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Revision of the southern South American species of Philonthus Stephens (Coleoptera: Staphylinidae)

Current affiliation: Laboratorio de Entomología, Instituto Argentino de Investigaciones de las Zonas Aridas (IADIZA, CCT CONICET, Mendoza), Casilla de Correo 507, 5500 Mendoza, Argentina Current address: Drusustrasse 22, (40549) Duesseldorf, Germany
Coleoptera Staphylininae Philonthina Philonthus Belonuchus taxonomy South America distribution

Abstract

The southern South American species of the genus Philonthus Stephens, 1829 (Staphylininae: Staphylinini) are revised using characters of external morphology and male and female genitalia. Three species of Philonthus are transferred to the genus Belonuchus Nordmann, 1837 resulting in the following new combinations: B. aluticollis (Bernhauer, 1921), B. flavicoxis (Bernhauer, 1912) and B. weiserianus (Bernhauer, 1921). Twenty-five valid species are recognized for the region in the present study: Philonthus aeruginosus Nordmann 1837, P. argus Herman 2001, P. bicoloristylus ChaniPosse 2004, P. bonariensis Bernhauer 1909, P. convexicollis Lynch-Arribálzaga 1884, P. cribriventris Bernhauer 1912, P. discoideus (Gravenhorst, 1802), P. figulus Erichson 1840, P. flavolimbatus Erichson 1840, P. hepaticus Erichson 1840, P. jenseni Bernhauer 1912, P. longicornis Stephens 1832, P. lynchi Bernhauer & Schubert 1914, P. pallipes Blanchard 1842, P. pauxillus Solsky 1868, P. politus (Linnaeus, 1758), P. quadraticeps Boheman 1858, P. rectangulus Sharp 1874, P. rubromaculatus Blanchard 1842, P. ruficauda Bernhauer 1934, P. sericans (Gravenhorst, 1802), P. stenocephalus Scheerpeltz 1972, P. varians (Paykull, 1789). Two new species are described: P. floresi and P. bruchianus. Philonthus feralis Erichson 1840, previously recorded for this region, is also redescribed. Diagnoses, redescriptions or descriptions, illustrations, bionomic information and distribution maps for all species are presented. An identification key to the southern South American species of the genus is provided. Lectotypes are designated for Philonthus aeruginosus Nordmann 1837, P. aluticollis Bernhauer 1921, P. apicipennis Lynch-Arribálzaga 1884, P. convexicollis LynchArribálzaga 1884, P. figulus Erichson 1840, P. flavicoxis Bernhauer 1912, P. jenseni Bernhauer 1912, P. pallipes Blanchard 1842, P. rubromaculatus Blanchard 1842, P. stenocephalus Scheerpeltz 1972, P. tucumanensis Bernhauer 1934 and P. weiserianus Bernhauer, 1921.

References

  1. Baudi di Selve, F. (1848) Alcune specie nuove di stafilini. Studi Entomologici, 1(2), 113–148.

    Bernhauer, M. (1909) Zur Staphylinidenfauna von Südamerika (6. Beitrag.). Bolletino della Società Entomologica Italiana, 40 (1908), 225–251.

    Bernhauer, M. (1912) Zur Staphylinidenfauna von Südamerika (10. Beitrag). Verhandlungen der k. k. zoologisch-Botanischen Gesellschaft in Wien, 62, 26–48.

    Bernhauer, M. (1916) Zur Staphylinidenfauna des tropischen Amerika, insbesondere der columbischen Cordilleren. Koleopterologische Rundschau, 5, 25–40.

    Bernhauer, M. (1921) Neue Staphyliniden aus Südamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101–108, 169–179.

    Bernhauer, M. (1927) Zur Staphylinidenfauna Süd-Amerikas, insbesondere Argentiniens. Archiv für Naturgeschichte, (A) 91(12) (1925), 229–264.

    Bernhauer, M. (1934) Neue Staphyliniden aus Argentinien (Col.) (34. Beitrag zur Kenntnis der südamerikanischen Fauna). Revista de Entomologia, 4, 112–119 (III).

    Bernhauer, M. & Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, 5(57). W. Junk, Berlin, pp. 289–408.

    Blackwelder, R.E. (1936) Morphology of the coleopterous family Staphylinidae. Smithsonian Miscelaneous Collection, 94(13), 1–102.

    Blackwelder, R.E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. United States National Museum Bulletin, 182, 1–658.

    Blackwelder, R.E. (1944) Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. Part I. United States National Museum Bulletin, 185, 1–188.

    Blanchard, C.E. (1842) “New species”. In: A. Brullé, Voyage dans l’Amérique méridionale . . . par Alcide d’ Orbigny . . .Insectes Coléoptères. 6(2), 57–88. P. Bertrand, Paris.

    Boheman, C.H. (1858) Coleoptera. Species novas descripsit. In: Virgin, C. (Ed.), Kongliga Svenska Fregatten Eugenies resa omkring Jorden, Zoologi I, Insecta. P. A. Norstedt & Söner, Stockholm, pp. 1–112.

    Boller, F. (1983) Zur Larvalmorphologie der Gattung Philonthus Curtis (Coleoptera: Staphylinidae). Spixiana, 6, 113–131.

    Bruch, C. (1915) Catálogo Sistemático de los coleópteros de la República Argentina. Pars VI. Revista del Museo de La Plata, 19 (Segunda Serie) (6), 235–302.

    Byrne, M. (1993) The immature stages of Philonthus sanamus Tottenham (Coleoptera: Staphylinidae). African Entomology, 1(2), 229–234.

    Cabrera-Walsh, G. & Cordo, H.A. (1997) Coprophilous arthropod community from Argentina species of potential use as biocontrol agents against pest flies. Environmental Entomology , 26, 191–200.

    Cabrera Walsh, G. & Chani Posse, M. (2003) Abundance and seasonal Distribution of predatory coprophilous Argentine rove beetles (Coleoptera: Staphylinidae), and their effects on dung breeding flies. Coleopterists Bulletin, 57(1), 43–50.

    Chani Posse, M.R. (2004) Eight Argentinean species of dung-inhabiting Philonthus Stephens (Coleoptera: Staphylinidae). Studies on Neotropical Fauna and Environment, 39, 217–232.

    Chani Posse, M. (2006a) Larval morphology and chaetotaxy of Philonthus Stephens (Coleoptera: Staphylinidae) based on descriptions of eight species from Argentina. Annales Zoologici, 56(1), 7–28.

    Chani Posse, M. (2006b) Systematic revision and cladistic analysis of the Neotropical genus Chroaptomus Sharp (Coleoptera: Staphylinidae), with descriptions of two new species. Insect Systematics & Evolution, 37, 361–383.

    Chani Posse de Maus, M. (2008) Systematic revision and cladistic analysis of the South American genus Endeius Coiffait & Sáiz (Coleoptera: Staphylinidae). Insect Systematics & Evolution, 39, 381–406.

    Chani Posse de Maus, M. (2009) Revision of some types of Philonthus Stephens and Gabrius Stephens from southern South America (Coleoptera: Staphylinidae), with nomenclatural notes. Zootaxa, 2034, 31–42.

    Chevrolat, L. & Fauvel, A. (1863) Coléoptères de l’Ile de Cuba. Notes, synonymiques et descriptions d’espèces nouvelles. Annales de la Société Entomologique de France, (4)3, 427–446.

    Coiffait, H. (1974) Coléoptères Staphylinides de la Région Paléarctique Occidentale. II. Sous famille Staphylininae, Tribus Philonthini et Staphylinini. Nouvelle Revue d´Entomologie, Supplément, 4(4), 1–593.

    Coiffait, H. & Sáiz, F. (1968) Les Staphylinidae (sensu lato) du Chili. In: Delamare Deboutteville, C. & Rapoport, E. (Eds.), Biologie de l’Amérique Australe. Études sur la faune du sol, 4. Centre National de la Recherche Scientifique, Paris, pp. 339–468.

    Curtis, J. (1829) A guide to an arrangement of British Insects: being a catalogue of all the named species hitherto discovered in Great Britain and Ireland. Westley, London, 6 pp., 248 columns.

    Curtis, J. (1836) British entomology; being illustrations and descriptions of the genera of insects found in Great Britain and Ireland: containing coloured figures from nature of the most rare and beautiful species, and in many instances of the plants upon which they are found. Volume XIII. Curtis, London, pls. 578–625.

    Dejean, P.F.M.A. (1821) Catalogue de la collection de Coléoptères de M. le Baron Dejean. Crevot Libraire, Paris, 136 pp.

    Desender, K. & Baert, L. (1996) The Coleoptera of Easter Island. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique. Entomologie, 66, 27–50.

    Eppelsheim, E. (1895) Beitrag zur Staphylinenfauna West-Afrika’s. (2. Stück.) Deutsche Entomologische Zeitschrift, 1895(1), 113–141.

    Erichson, W.F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, pp 401–954.

    Fabricius, J.C. (1801) Systema eleutheratorum secundum ordines, genera, species: adiectis synonymis, locis, observationibus, descriptionibus. Vol. 2. Bibliopolii Academici Novi, Kiliae, 687 pp.

    Fairmaire, L. & Germain, P. (1861) Révision des Coléoptères du Chili (suite). Annales de la Société Entomologique de France, (4)1, 405–456.

    Fauvel, A. (1866a) Faune du Chili. Insectes Coléoptères. Staphylinides. Bulletin de la Société Linnéenne de Normandie, 10, 250–353. “Note: Same as Fauvel, 1866b”

    Fauvel, A. (1866b) Faune du Chili. Insectes Coléoptères. Staphylinides. Notices Entomologiques, 4, 1–103. “Note: Same as Fauvel, 1866a”

    Fauvel, A. (1869) Remarques synonymiques sur les Staphylinidae du Catalogus Coleopterorum de M. M. Harold et Gemminger. L’Abeille. Memoires d’Entomologie, 5, 479–94.

    Fauvel, A. (1874a) Faune Gallo-Rhénane ou species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la Prusse Rhénane, le Nassau et le Valais avec tableaux synoptiques et planches gravées. Coléoptères. Bulletin de la Société Linnéene de Normandie, (2)8, 167–340.

    Fauvel, A. (1874b) Faune Gallo-Rhénane ou species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la Prusse Rhénane, le Nassau et le Valais avec tableaux synoptiques et planches gravées. Coléoptères. Vol. 3, liv. 5. F. le Blanc-Hardel, Caen, pp. 391–544. “Note: Same as Fauvel, 1874a”

    Fauvel, A. (1877) Les staphylinides de l’Australie et de la Polynésie. Annali del Museo

    Civico di Storia Naturale di Genova, 10, 168–298.

    Fauvel, A. (1891) Voyage de M. E. Simon au Vénézuela (Décembre 1887-Avril 1888). 11e Mémoire. Revue d´Entomologie, 10, 87–127.

    Fauvel, A. (1903) Faune analytique des coléoptères de la Nouvelle Calédonie. Revue d'Entomologie (Caen), 22, 203–379.

    Fincher, G.T & Summerlin, J.W. (1994) Predation on the horn fly by three exotic species of Philonthus. Journal of Agricultural Entomology, 11, 1, 45–48.

    Flechtmann, C.A.H., Rodrigues, S.R. & Gaspareto, C.L. (1995a) Controle biológico da mosca-dos-chifres (Haematobia irritans irritans) em Selvíria, Mato Grosso do Sul. 6. Dinâmica populacional de insetos fimícolas excetuando-se Scarabaeidae coprófagos. Revista Brasileira de Entomolgia, 39, 287–296.

    Flechtmann, C.A.H., Rodrigues, S.R., de Araújo, S.D. & Wenzel, R.L. (1995b) Levantamento de insetos fimícolas em Ilha Solteira, São Paulo, Brasil. Revista Brasileira de Entomolgia, 39, 115–120.

    Frank, J.H. (1982) The parasites of the Staphylinidae (Coleoptera). A contribution towards an encyclopedia of the Staphylinidae. University of Florida Agr. Exp. Stations, Technical Bulletin, 824, i–vii +1–118 pp.

    Ganglbauer, L. (1895) Die Käfer von Mitteleuropa. Die Käfer der österreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des französischen und italienischen Alpengebietes. 2. Familienreihe Staphylinoidea. Theil I. Staphylinidae, Pselaphidae. Carl Gerold's Sohn, Wien, 881 pp.

    Gravenhorst, J.L.C. (1802) Coleoptera Microptera Brunsvicencia nec non exoticorum quotquot exstant in collectionibus entomologorum brunvicencsium in genera familias et species distribuit. Carolus Reichard, Brunsvigae, 206 pp.

    Gridelli, E. (1920) Terzo contributo alla conoscenza delle specie paleartiche del genere Philonthus Steph. Atti dell´Accademia Scientifica veneto Trentino Istriana, 11, 15–21.

    Guérin‑Méneville, M.F.E. (1855) Catalogue des insectes coléopteres, recueillis par M. Gaetano Osculati, pendant son exploration de la région équatoriale, sur les bords du Napo et de l'Amazone. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 5, 573–612.

    Guimarães, J.A. & Mendes, J. (1998) Succession and abundance of Staphylinidae in cattle dung in Uberlândia, Brazil. Memórias do Instituto Oswaldo Cruz, 93, 127–131.

    Harris, R.L. & Oliver, L.M. (1979) Predation of Philonthus flavolimbatus on the Horn Fly. Environmental Entomology, 8, 259–260.

    Hatch, M. & Ortenburger, A.I. (1930) Records and new species of Coleoptera from Oklahoma. Publications of the University of Oklahoma Biological Survey, 2(1), 7–14.

    Herman, L.H. (2001a) Nomenclatural changes in the Staphylinidae (Insecta: Coleoptera). Bulletin of the American Museum of Natural History, 264, 1–83.

    Herman, L.H. (2001b) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441–3020.

    Hochhuth, J.H. (1849) Die Staphylinen-Fauna des Kaukasus und Transkaukasiens. Bulletin de la Société Impériale des Naturalistes de Moscou, 22(1), 18–214.

    Holme, F. (1837) Notice of the coleopterous insects observed in the Scilly Islands in July and August, 1836. The Transactions of the Entomological Society of London, 2(1), 58–64.

    Horn, W., Kahle I., Friese G., & Gaedike R. (1990) Collectiones Entomologicae. Akademie der Landwirtschaftswissenschaften der Deutschen Demokratischen Republik, 2 parts, 573 pp.

    Hu, G.Y. & Frank, J.H. (1995) Structural comparison of the chorion surface of five Philonthus species (Coleoptera: Staphylinid). Proceedings of the Entomological Society of Washington, 97(3), 582–589.

    Hu, G.Y. & Frank, J.H. (1997) Predation on the horn fly (Diptera: Muscidae) by five species of Philonthus (Coleoptera: Staphylinidae). Environmental Entomology, 26, 1240–1246.

    Hunter III, J.S., Bay, D.E., & Fincher, G.T. (1986) A Survey of Staphylinidae associated with cattle droppings in Burleson County, Texas. Southwestern Entomology, 11(2), 83–87.

    Hunter III, J.S., Fincher, G.T., Bay, D.E. & Beerwinkle, K.R. (1991) Seasonal distribution and diel flight activity of Staphylinidae (Coleoptera) in open and wooded pasture in east-central Texas. Journal of the Kansas Entomological Society, 64(2), 163–173.

    ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp.

    Kasule, F.K. (1970) The larvae of Paederinae and Staphylininae (Coleoptera: Staphylinidae) with keys to the known British genera. Transactions of the Royal Entomological Society of London, 122(2), 49–80.

    Koller, W.W., Gomes, A., Rodrigues, S.R. & Mendes, J. (2002) Staphylinidae (Coleoptera) Associated to Cattle Dung in Campo Grande, MS, Brazil. Neotropical Entomology [online], 31(4), 641–645.

    Kraatz, G. (1859) Zur kritischen Kenntniss der in Gay’s Historia Fisica y Politica von Solier beschriebenen Staphylinen. Berliner Entomologische Zeitschrift, 3, 1–16.

    Kranebitter, P. & Schatz, I. (2002) The larval instars of Philonthus aerosus Kiesenwetter (subgenus Kenonthus Coiffait) from the central Alps (Tyrol, Austria) with a comparison of the known species of the genus (Coleoptera, Staphylinidae). Entomologische Blätter, 98, 61–64.

    Linnaeus, C. v. (1758) Systema naturae, per regna tria naturae, secundum Classes, Ordines, Genera, Species, cum characteribus, differentiis, synonymis, locis. vol. 1. ed. 10. Laurentii Salvii, Holmiae “= Stockholm”, 824 pp.

    Lynch-Arribálzaga, F. (1884) Estafilinos de Buenos Aires. Boletín de la Academia Nacional de Ciencias, Córdoba, 7, 5–256.

    Morrone, J.J. (2006) Biogeographic areas and transition zones of Latin America and the Caribbean islands based on panbiogeographic and cladistic analyses of the entomofauna. Annual Review of Entomology, 51, 467–494.

    Navarrete-Heredia, J.L., Newton, A.F., Thayer, M.K., Ashe, J.S. & Chandler, D.S. (2002) Guía ilustrada para los géneros de Staphylinidae (Coleoptera) de México. Illustrated guide to the genera of Staphylinidae (Coleoptera) of México. Universidad de Guadalajara y Conabio, México. 401 pp.

    Newton, A. F. & Thayer, M. K. (2005) Catalog of higher taxa of Staphyliniformia and genera and subgenera of Staphylinoidea, Field Museum of Natural History, Chicago. Available from: http://www.fieldmuseum.org/peet_staph/db_1a.html (last updated 03 November 2005; date of access: 17 December 2008).

    Nordmann, A. von. (1837) Symbolae ad monographiam staphylinorum. Ex Academiae Caesareae Scientiarum Tomo IV, Petropoli, 167 pp., 2 pls.

    Paykull, G. de (1789) Monographia Staphylinorum Sueciae. Johann Edman, Upsaliae, 8 + 81 pp.

    Pietrykowska-Tudruj, E. & Staniec, B. (2006) Description of the egg and larva of Philonthus punctus (Gravenhorst, 1802) (Coleoptera, Staphylinidae, Staphylininae). Deutsche Entomologische Zeitschrift, 53 (2), 179–192.

    Pietrykowska-Tudruj, E. & Staniec, B. (2008) Morphology of developmental stages of Philonthus fumarius (Gravenhorst, 1806) (Coleoptera, Staphylinidae) with notes on biology. Acta Zoologica Academiae Scientiarum Hungaricae, 54 (3), 213–234.

    Pototskaya, V.A. (1966) Les larves de la tribe Philonthini (Coleoptera: Staphylinidae). Revue d´Ecologie et de Biologie du Sol 3, 141–162.

    Rambousek, F. (1925) Staphylinides nouveaux de l’Amérique meridionale. (2e contribution). Časopis Československé Společnosti Entomologické, 21, 69–76.

    Roth, J.P. (1982) Predation on the horn fly, Haematobia irritans (L.), by three Philontus species. Southwestern Entomology, 7(1), 26–30.

    Sáiz, F. (1971) Philonthini y Staphylinini de Chile (Coleoptera: Staphylinidae). Boletín del Museo Nacional de Historia Natural, Chile, 29, 339–353.

    Sahlberg, R.F. (1847) Coleoptera diebus XV–XXVII Decembris anni MDCCCXXXIX ad Rio Janeiro lecta. Acta Societatis Scientiarum Fennicae, 2(2), 787–805.

    Say, T. (1830) Description of new species of North American insects, and observations on some of the species already described. School Press, New Harmony (IN), pp. 1–41.

    Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 129. W. Junk, Berlin, pp. 989–1500.

    Scheerpeltz, O. (1960) Zur Kenntnis neotropischer Staphyliniden (Col.). Beiträge zur Neotropischen Fauna, 2, 65–138.

    Scheerpeltz, O. (1972) Wissenschaftliche Ergebnisse der Studienreise von Gy. Topál nach Südwest-Argentinien (Coleoptera: Staphylinidae). Folia Entomologica Hungarica (N.S.) 25 (Suppl.), 268 pp., 5 pls.

    Schillhammer, H. (1997) Taxonomic revision of the Oriental species of Gabrius Stephens. Monographs on Coleoptera, 1, 1–139.

    Schillhammer, H. (1998) Revision of the East Palaearctic and Oriental species of Philonthus Stephens-Part 1. The cyanipennis group. Koleopterologische Rundschau, 68, 101–118.

    Schillhammer, H. (1999a) Revision of the East Palaearctic and Oriental species of Philonthus Stephens-Part 2. The spinipes and cinctulus groups. Koleopterologische Rundschau, 69, 55–65.

    Schillhammer, H. (1999b) Synopsis of the genus Hesperopalpus Shibata, 1973 with description of a new species from Vietnam. Annalen des Naturhistorischen Museums, 101 B, 303–307.

    Schillhammer, H. (2000a) Revision of the East Palaearctic and Oriental species of Philonthus Stephens-Part 3. The politus complex. Koleopterologische Rundschau, 70, 113–176.

    Schillhammer, H. (2000b) A new genus and species of the subtribe Philonthina from Kashmir (Coleoptera: Staphylinidae). Entomological Problems, 31 (1), 35–37.

    Schillhammer, H. (2000c) New species and records of Oriental and East Palaearctic species of Gabrius Stephens, 1829. Annalen des Naturhistorischen Museums, 102 B, 117–142.

    Schillhammer, H. (2001a) Revision of the East Palaearctic and Oriental species of Philonthus Stephens-Part 4. The amplitarsis group and an additional species of the cinctulus group. Koleopterologische Rundschau, 71, 59–65.

    Schillhammer, H. (2001b) The Taiwan species of Gabrius Stephens. Annalen des Naturhistorischen Museums, 103 B, 391–411.

    Schillhammer, H. (2002) Three new species of Hesperus Fauvel from South-east Asia. Koleopterologische Rundschau, 72, 127–135.

    Schillhammer, H. (2003a) On some Central Asian species of the Gabrius astutus group. Annalen des Naturhistorischen Museums, 104 B, 353–361.

    Schillhammer, H. (2003b) Revision of the East Palaearctic and Oriental species of Philonthus Stephens-Part 5. The rotundicollis and sanguinolentus species groups. Koleopterologische Rundschau, 73, 85–136.

    Sharp, D.S. (1876) Contributions to an insect fauna of the AmazonValley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27–424.

    Sharp, D.S. (1885) Staphylinidae. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. 1(2). Taylor & Francis, London, pp. 393–536.

    Smetana, A. (1958) Fauna ČSR. Svazek 12. Drabčikoviti – Staphylinidae. I. Staphylininae: (Řád: Brouci – Coleoptera). Československé Akademie Věd, Praha, 435 pp.

    Smetana, A. (1982) Revision of the subfamily Xantholininae of America North of Mexico (Coleoptera: Staphylinidae). Memories of the Entomological Society of Canada, 12, 1–389.

    Smetana, A. (1994) Lectotype designations and redescriptions of three New World species of Philonthus Stephens, 1829, described by Erichson in 1840 (Coleoptera Staphylinidae). Nouvelle Revue d’Entomologie, (n. ser.) 10(1993): 341–348.

    Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1–946.

    Smetana, A. & Davies, A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 1–88.

    Solier, A.J.J. (1849) Orden III. Coleópteros. In: Gay, C. (ed.), Historia Física y Política de Chile. Zoología, Vol. 4. Imprenta de Maulde et Renou, Paris, pp. 105–380.

    Solsky, S. (1868) Études sur les staphylinids du Mexique. Horae Societatis Entomologicae Rossicae (St. Petersburg), 5, 119–144.

    Staniec, B. & Pietrykowska, E. (2005) The pupae of Gyrohypnus fracticornis (Mueller, 1776) and Philonthus tenuicornis Mulsant & Rey, 1853 (Coleoptera: Staphylinidae). Genus, 16(3), 331–339.

    Staniec, B. & Pietrykowska-Tudruj, E. (2007) Developmental stages of Philonthus rubripennis Stephens, 1832 (Coleoptera, Staphylinidae, Staphylininae) with comments of its biology. Deutsche Entomologische Zeitschrift, 54 (1), 95–113.

    Steinheil, E. (1869) Symbolae ad historiam coleopterorum Argentiniae meridionalis, . . . Atti della Società Italiana di Scienze Naturali, 12, 238–260.

    Stephens, J.F. (1829) The Nomenclature of British Insects; Being a Compendious List of Such Species as are Contained in the Systematic Catalogue of British Insects, and Forming a Guide to their Classification. Baldwin & Cradock, London, 68 pp.

    Stephens, J.F. (1832) Illustrations of British Entomology, or, a Synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. 5. Baldwin & Cradock, London, 447 pp.

    Waterhouse, G.R. (1858) Catalog of British Coleoptera. Taylor and Francis, London, iv + 3–117 pp.