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Issue: Vol. 1 No. 3: In progress
Type: Article
Published: 2024-08-05
DOI: 10.11646/bpn.1.3.2
Page range: 44-82
Abstract views: 543
PDF downloaded: 185

Phylogenetic nomenclature of Notosuchia (Crocodylomorpha; Crocodyliformes)

CONICET, Instituto de Estudios Andinos “Don Pablo Groeber” (IDEAN), Departamento de Ciencias Geológicas, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Intendente Güiraldez 2160, Ciudad Universitaria, Pabellón 2, C1428EGBA, Buenos Aires, Argentina; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Intendente Güiraldez 2160, Ciudad Universitaria, Pabellón 2, C1428EGBA, Buenos Aires, Argentina.
CONICET, Museo Argentino Ciencias Naturales “Bernardino Rivadavia“, Avenida Ángel Gallardo 470, CP 1405, Buenos Aires, Argentina.
Laboratório de Paleontologia e Evolução de Ilha Solteira, Universidade Estadual Paulista (UNESP), Rua Monção 226, 15385000, Ilha Solteira, São Paulo, Brazil.
Centro de Pesquisas Paleontológicas “Llewellyn Ivor Price”, Complexo Cultural e Científico Peirópolis, Pró-Reitoria de Extensão Universitária, Universidade Federal do Triângulo Mineiro, Uberaba, Minas Gerais, Brazil; Departamento de Ciências Biológicas, Instituto de Ciências Exatas, Naturais e Educação (ICENE), Universidade Federal do Triângulo Mineiro (UFTM), Av. Randolfo Borges Jr. 1400, Univerdecidade, Uberaba, Minas Gerais, 38064-200, Brazil.
Laboratório de Paleontologia e Evolução de Ilha Solteira, Universidade Estadual Paulista (UNESP), Rua Monção 226, 15385000, Ilha Solteira, São Paulo, Brazil; Programa de Pós Graduação em Biodiversidade, Instituto de Biociências, Letras e Ciências Exatas, Universidade Estadual Paulista (UNESP), R. Cristóvão Colombo, 2265, 15054-000, São José do Rio Preto, São Paulo, Brazil; Senckenberg Centre for Human Evolution and Paleoenvironment, Eberhard Karls University of Tübingen, Sigwartstraße 10, 72070, Tübingen, Germany.
CONICET, Instituto Superior de Correlación Geológica, Universidad Nacional de Tucumán, Facultad de Ciencias Naturales e I.M.L., Av. Presidente Perón S/N, Yerba Buena, 4107, Tucumán, Argentina.
Departamento de Ciências, Faculdade de Formação de Professores, Universidade do Estado do Rio de Janeiro (DCIEN/FFP/UERJ), Francisco Portela, 1470—Patronato, São Gonçalo city, 24435-005, Rio de Janeiro, Brazil.
Department of Zoology, Institute of Biosciences, University of São Paulo, São Paulo, Brazil.
Department of Earth Sciences, University College London, Gower Street, London, WC1E 6BT, United Kingdom.
Universidad Nacional de Río Negro, Instituto de Investigación en Paleobiología y Geología, Río Negro, Argentina; CONICET, Instituto de Investigación en Paleobiología y Geología (IIPG), Av. Roca 1242, R8332EXY, General Roca, Río Negro, Argentina.
Redpath Museum, McGill University, 859 Sherbrooke St. W., Montreal, Quebec H3A 0C4, Canada.
Ziphosuchia Sebecia Xenodontosuchia Peirosauria Sphagesauria definition registration

Abstract

Notosuchia is a large and diverse clade of Mesozoic crocodyliforms that thrived in continental environments especially during the Cretaceous of Gondwana. This clade has been the focus of several phylogenetic studies that led to the consensus of two main topological arrangements, mostly differing in the position of the clade Sebecidae: the sebecosuchian and sebecian hypotheses. In this manuscript we review the main phylogenetic definitions of several clades of notosuchians in the context of these hypotheses. As a result, besides the clade names previously established under the ICPN/Phylocode (including registration in RegNum) by Ruiz et al. (2021) (Notosuchia, Xenodontosuchia, Sphagesauria, Sphagesauridae) and Darlim et al. (2021a) (Baurusuchia, Baurusuchidae, Baurusuchinae, Pissarrachampsinae), eleven clade names have been registered and are defined in this contribution. Ten of these represent converted clade names (Uruguaysuchidae, Mahajangasuchidae, Peirosauridae, Itasuchidae, Ziphosuchia, Sphagesaurinae, Caipirasuchinae, Sebecia, Sebecosuchia, and Sebecia), while one represents a new clade name (Peirosauria). Furthermore, the definition of Xenodontosuchia is emended (unrestricted emendation) so that the name can be used in the context of a wider array of topologies.

 

References

  1. Andrade, M. B., and R. J. Bertini. 2008. A new Sphagesaurus (Mesoeucrocodylia: Notosuchia) from the Upper Cretaceous of Monte Alto city (Bauru Group, Brazil), and a revision of the Sphagesauridae. Historical Biology 20(2):101–136. https://doi.org/10.1080/08912960701642949
  2. Andrade, M. B., R. Edmonds, M. J. Benton, and R. Schouten. 2011. A new Berriasian species of Goniopholis (Mesoeucrocodylia, Neosuchia) from England, and a review of the genus. Zoological Journal of the Linnean Society 163(1):S66–S108. https://doi.org/10.1111/j.1096-3642.2011.00709.x
  3. Antunes, M. T. 1975. Iberosuchus, crocodile sebecosuchian nouveau, l’Eocène iberique au Nord de la Chaîne Centrale, et l’origine du canyon de Nazaré. Comunicações Servicio Geologico Portugal 49:285–330.
  4. Barrios, F., A. Paulina-Carabajal, and P. Bona. 2016. A new peirosaurid (Crocodyliformes, Mesoeucrocodylia) from the Upper Cretaceous of Patagonia, Argentina. Ameghiniana 53:14–25. https://doi.org/10.5710/AMGH.03.09.2015.2903
  5. Benton, M. J., and J. M. Clark. 1988. Archosaur phylogeny and the relationships of the Crocodylia. Pp. 295–338 in The Phylogeny and Classification of the Tetrapods (M. J. Benton, ed.). Clarendon Press, Oxford.
  6. Berg, D. 1966. Die Krokodile, insbesondere Asiatosuchus und aff. Sebecus? aus dem Eozän von Messel bei Darmstadt/Hessen. Abhandlungen des Hessischen Landesamtes für Bodenforschung 52:1–105.
  7. Bravo, G. G., D. Pol, and D. A. García-López. 2021. A new sebecid mesoeucrocodylian from the Paleocene of Northwestern Argentina. Journal of Vertebrate Paleontology 41(3):e1979020. https://doi.org/10.1080/02724634.2021.1979020
  8. Buckley, G. A., and C. A. Brochu. 1999. An enigmatic new crocodile from the Upper Cretaceous of Madagascar. Special Papers in Palaeontology 60:149–175. https://doi.org/10.1080/02724634.2021.1979020
  9. Buckley, G. A., C. A. Brochu, D. W. Krause, and D. Pol. 2000. A pugnosed crocodyliform from the Late Cretaceous of Madagascar. Nature 405:941–944. https://doi.org/10.1038/35016061
  10. Buffetaut, E. 1989. A new ziphodont mesosuchian crocodile from the Eocene of Algeria. Palaeontographica 208:1–10.
  11. Cantino, P. D., and K. de Queiroz. 2020. International Code of Phylogenetic Nomenclature (PhyloCode). CRC Press, Boca Raton, Florida. https://doi.org/10.1201/9780429446320
  12. Carvalho, I. S., L. C. B. Ribeiro, and L. dS. Avila. 2004. Uberabasuchus terrificus sp. nov., a new crocodylomorph from the Bauru Basin (Upper Cretaceous), Brazil. Gondwana Research 7(4):975–1002. https://doi.org/10.1016/S1342-937X(05)71079-0
  13. de Celis, A., I. Narváez, A. Arcucci, and F. Ortega. 2021. Lagerstätte effect drives notosuchian palaeodiversity (Crocodyliformes, Notosuchia). Historical Biology 33(11):3031–3040. https://doi.org/10.1080/08912963.2020.1844682
  14. Clark, J. M. 1994. Patterns of evolution in Mesozoic Crocodyliformes. Pp. 84–97 in In the Shadow of the Dinosaurs, Early Mesozoic Tetrapods (N. C. Fraser, and H.-D. Sues, eds.). Cambridge University Press, Cambridge.
  15. Colbert, E. H. 1946. Sebecus, representative of a peculiar suborder of fossil Crocodilia from Patagonia. Bulletin of the American Museum of Natural History 87:217–270.
  16. Company, J., X. Pereda-Suberbiola, J. I. Ruíz-Omeñaca, and A. D. Buscalioni. 2005. A new species of Doratodon (Crocodyliformes: Ziphosuchia) from the Late Cretaceous of Spain. Journal of Vertebrate Paleontology 25(2): 343–353. https://doi.org/10.1671/0272-4634(2005)025[0343:ANSODC]2.0.CO;2
  17. Darlim, G., F. C. Montefeltro, and M. C. Langer. 2021a. 3D skull modelling and description of a new baurusuchid (Crocodyliformes, Mesoeucrocodylia) from the Late Cretaceous (Bauru Basin) of Brazil. Journal of Anatomy 239(3):622–662. https://doi.org/10.1111/joa.13442
  18. Darlim, G., I. S. Carvalho, S. A. S. Tavares, and M. C. Langer. 2021b. A new Pissarrachampsinae specimen from the Bauru Basin, Brazil, adds data to the understanding of the Baurusuchidae (Mesoeucrocodylia, Notosuchia) distribution in the Late Cretaceous of South America. Cretaceous Research 128:104969. https://doi.org/10.1016/j.cretres.2021.104969
  19. Fernández Dumont, M. L., P. Bona, D. Pol, and S. Apesteguía. 2020. New anatomical information on Araripesuchus buitreraensis with implications for the systematics of Uruguaysuchidae (Crocodyliforms, Notosuchia). Cretaceous Research 113:104494. https://doi.org/10.1016/j.cretres.2020.104494
  20. Gasparini, Z. 1971. Los Notosuchia del Cretácico de América del Sur como un nuevo Infraorden de los Mesosuchia (Crocodilia). Ameghiniana 8:83–103.
  21. Gasparini, Z. 1972. Los Sebecosuchia (Crocodilia) del Territorio Argentino. Consideraciones sobre su “status” taxonómico. Ameghiniana 9:23–34.
  22. Gasparini, Z. 1982. Una nueva familia de cocodrilos zifodontes cretácicos de América del Sur. Actas V Congreso Latinoamericano de Geología, Buenos Aires, 1981 4:317–329.
  23. Gasparini, Z., L. M. Chiappe, and M. Fernández. 1991. A new senonian peirosaurid (Crocodylomorpha) from Argentina and a synopsis of the South American Cretaceous crocodilians. Journal of Vertebrate Paleontology 11(3):316–333. https://doi.org/10.1080/02724634.1991.10011401
  24. Gasparini, Z., D. Pol, and L. A. Spalletti. 2006. An unusual marine crocodyliform from the Jurassic-Cretaceous boundary of Patagonia. Science 311:70–72. https://doi.org/10.1126/science.1120803
  25. Geroto, C. F. C., and R. J. Bertini. 2019. New material of Pepesuchus (Crocodyliformes; Mesoeucrocodylia) from the Bauru Group: implications about its phylogeny and the age of the Adamantina Formation. Zoological Journal of the Linnean Society 185:312–334. https://doi.org/10.1093/zoolinnean/zly037
  26. Godoy, P. L., F. C. Montefeltro, M. A. Norell, and M. C. Langer. 2014. An additional baurusuchid from the Cretaceous of Brazil with evidence of interspecific predation among Crocodyliformes. PLoS ONE 9(5):e97138. https://doi.org/10.1371/journal.pone.0097138
  27. Godoy, P. L., M. Bronzati, E. Eltink, J. C. A. Marsola, G. M. Cidade, M. C. Langer, and F. C. Montefeltro. 2016. Postcranial anatomy of Pissarrachampsa sera (Crocodyliformes, Baurusuchidae) from the Late Cretaceous of Brazil: Insights on lifestyle and phylogenetic significance. PeerJ 4:e2075. https://doi.org/10.7717/peerj.2075
  28. Gomani, E. M. 1997. A crocodyliform from the Early Cretaceous dinosaur beds, northern Malawi. Journal of Vertebrate Paleontology 17(2):280–294. https://doi.org/10.1080/02724634.1997.10010975
  29. Iori, F. V., and I. S. Carvalho. 2011. Caipirasuchus paulistanus, a new sphagesaurid (Crocodylomorpha, Mesoeucrocodylia) from the Adamantina Formation (Upper Cretaceous, Turonian–Santonian), Bauru Basin, Brazil. Journal of Vertebrate Paleontology 31(6):1255–1264. https://doi.org/10.1080/02724634.2011.602777
  30. Iori, F. V., T. S. Marinho, I. S. Carvalho, and A. C. A. Campos. 2011. Padrão Dentário dos Esfagessaurídeos (Crocodyliformes, Sphagesauridae). Pp. 583–592 in Paleontologia: Cenários de Vida, v. 4 (I. S. Carvalho, N. K. Srivastava, O. Strohschoen Jr., C. Cunha Lana Org.). Editora Interciência, Rio de Janeiro.
  31. Iori, F. V., T. S. Marinho, I. S. Carvalho, and A. C. A. Campos. 2013. Taxonomic reappraisal of the sphagesaurid crocodyliform Sphagesaurus montealtensis from the Late Cretaceous Adamantina Formation of São Paulo State, Brazil. Zootaxa 3686(2):183–200. https://doi.org/10.11646/zootaxa.3686.2.4
  32. Iori, F. V., T. S. Marinho, I. S. Carvalho, and L. A. S. Frare. 2018. Cranial morphology of Morrinhosuchus luziae (Crocodyliformes, Notosuchia) from the Upper Cretaceous of the Bauru Basin, Brazil. Cretaceous Research 86:41e52. https://doi.org/10.1016/j.cretres.2018.02.010
  33. Kellner, A. W. A., A. E. P. Pinheiro, and D. A. Campos. 2014. A new sebecid from the Paleogene of Brazil and the crocodyliform radiation after the K–Pg boundary. PLoS ONE 9:e81386. https://doi.org/10.1371/journal.pone.0081386
  34. Kuhn, O. 1968. Die vorzeitlichen Krokodile. Ocben Krailling, München.
  35. Larsson, H. C. E., and H.-D. Sues. 2007. Cranial osteology and phylogenetic relationships of Hamadasuchus rebouli (Crocodyliformes: Mesoeucrocodylia) from the Cretaceous of Morocco. Zoological Journal of the Linnean Society 149:533–567. https://doi.org/10.1111/j.1096-3642.2007.00271.x
  36. Leardi, J. M., and D. Pol. 2009. The first crocodyliform from the Chubut Group (Chubut Province, Argentina) and its phylogenetic position within basal Mesoeucrocodylia. Cretaceous Research 30:1376–1386. https://doi.org/10.1016/j.cretres.2009.08.002
  37. Leardi, J. M., D. Pol, F. E. Novas, and M. Suárez Riglos. 2015. The postcranial anatomy of Yacarerani boliviensis and the phylogenetic significance of the notosuchian postcranial skeleton. Journal of Vertebrate Paleontology 35:e995187. https://doi.org/10.1080/02724634.2014.995187
  38. Leardi, J. M., D. Pol, and Z. Gasparini. 2018. New Patagonian baurusuchids (Crocodylormorpha; Notosuchia) from the Bajo de la Carpa Formation (Upper Cretaceous; Neuquén, Argentina): New evidences of the early sebecosuchian diversification in Gondwana. Comptes Rendus Paleovol 17:504–521. https://doi.org/10.1016/j.crpv.2018.02.002
  39. Lio, G., F. L. Agnolín, R. Juarez Valeri, L. Filippi, and D. Rosales. 2016. A new peirosaurid (Crocodyliformes) from the Late Cretaceous (Turonian-Coniacian) of Patagonia, Argentina. Historical Biology 28:835–841. https://doi.org/10.1080/08912963.2015.1043999
  40. Maganuco, S., C. Dal Sasso, and G. Pasini. 2006. A new large predatory archosaur from the Middle Jurassic (Bathonian) of Madagascar. Atti della Società italiana di scienze naturali e del museo civico di storia naturale di Milano 147(1):19.
  41. Marinho, T. S., and I. S. Carvalho. 2009. An armadillo-like sphagesaurid crocodyliform from the Late Cretaceous of Brazil. Journal of South American Sciences 27:36–41. https://doi.org/10.1016/j.jsames.2008.11.005
  42. Marinho, T. S., F. V. Iori, I. S. Carvalho, and F. M. Vasconcellos. 2013. Gondwanasuchus scabrosus gen. et sp. nov., a new terrestrial predatory crocodyliform (Mesoeucrocodylia: Baurusuchidae) from the Late Cretaceous Bauru Basin of Brazil. Cretaceous Research 44:104–111. https://doi.org/10.1016/j.jsames.2008.11.005
  43. Marinho, T. S., A. G. Martinelli, G. Basilici, M. V. T. Soares, A. Marconato, L. C. B. Ribeiro, and F. V. Iori. 2022. First Upper Cretaceous notosuchians (Crocodyliformes) from the Uberaba Formation (Bauru Group), southeastern Brazil: Enhancing crocodyliform diversity. Cretaceous Research 129:105000. https://doi.org/10.1016/j.cretres.2021.105000.
  44. Martin, J. E., and F. de Broin. 2016. A miniature notosuchian with multicuspid teeth from the Cretaceous of Morocco. Journal of Vertebrate Paleontology 36(6):e1211534. https://doi.org/10.1080/02724634.2016.1211534
  45. Martin, J. E., Y. Pochat-Cottilloux, Y. Laurent, V. Perrier, E. Robert, and P.-O. Antoine. 2023. Anatomy and phylogeny of an exceptionally large sebecid (Crocodylomorpha) from the middle Eocene of southern France. Journal of Vertebrate Paleontology 42:e2193828. https://doi.org/10.1080/02724634.2023.2193828
  46. Martinelli, A. G. 2003. New cranial remains of the bizarre notosuchid Comahuesuchus brachybuccalis (Archosauria, Crocodyliformes) from the Late Cretaceous of Río Negro Province (Argentina). Ameghiniana 40:559–572.
  47. Martinelli, A. G., J. J. W. Sertich, A. C. Garrido, and A. M. Praderio. 2012. A new peirosaurid from the Late Cretaceous of Argentina: implications for specimens referred to Peirosaurus torminni Price (Crocodyliformes: Peirosauridae). Cretaceous Research 37:191–200. https://doi.org/10.1016/j.cretres.2012.03.017
  48. Martinelli, A. G., T. S. Marinho, F. V. Iori, and L. C. B. Ribeiro. 2018. The first Caipirasuchus (Mesoeucrocodylia, Notosuchia) from the Late Cretaceous of Minas Gerais, Brazil: new insights on sphagesaurid anatomy and taxonomy. PeerJ 6:e5594. https://doi.org/10.7717/peerj.5594
  49. Martins, K. C., M. V. Lopes Queiroz, J. V. Ruiz, M. C. Langer, and F. C. Montefeltro. 2023. A new Baurusuchidae (Notosuchia, Crocodyliformes) from the Adamantina Formation (Bauru Group, Upper Cretaceous), with a revised phylogenetic analysis of Baurusuchia. Cretaceous Research 153:105680. https://doi.org/10.1016/j.cretres.2023.105680.
  50. Meunier, L. M. V., and H. C. Larsson. 2018. Trematochampsa taqueti as a nomen dubium and the crocodyliform diversity of the Upper Cretaceous in Beceten Formation of Niger. Zoological Journal of the Linnean Society 182(3):659–680. https://doi.org/10.1093/zoolinnean/zlx061
  51. Molnar, R. 1978. The crocodile from Tea Tree Cave and ziphodont crocodiles in Australia. Journal of Sydney Speleological Society 22(1):3–10.
  52. Montefeltro, F. C., H. C. Larsson, and M. C. Langer. 2011. A new baurusuchid (Crocodyliformes, Mesoeucrocodylia) from the Late Cretaceous of Brazil and the phylogeny of Baurusuchidae. PLoS One 6:e21916. https://doi.org/10.1371/journal.pone.0021916
  53. Montefeltro, F. C., H. C. Larsson, M. A. G. de França, and M. C. Langer. 2013. A new neosuchian with Asian affinities from the Jurassic of northwestern Brazil. Naturwissenschaften 100:835–841. https://doi.org/10.1007/s00114-013-1083-9
  54. Nascimento, P. M., and H. Zaher. 2010. A new species of Baurusuchus (Crocodyliformes, Mesoeucrocodylia) from the Upper Cretaceous of Brazil, with the first complete postcranial skeleton described for the family Baurusuchidae. Papéis Avulsos de Zoologia 50:323–361. https://doi.org/10.1590/S0031-10492010002100001
  55. Nicholl, C. S. C., E. S. E. Hunt, D. Ouarhache, and P. D. Mannion. 2021. A second peirosaurid crocodyliform from the Mid-Cretaceous Kem Kem Group of Morocco and the diversity of Gondwanan notosuchians outside South America. Royal Society Open Science 8:211254. https://doi.org/10.1098/rsos.211254
  56. O’Connor, P. M., J. J. Sertich, N. J. Stevens, E. M. Roberts, M. D. Gottfried, T. L. Hieronymus, and J. Temba. 2010. The evolution of mammal-like crocodyliforms in the Cretaceous Period of Gondwana. Nature 466:748–751. https://doi.org/10.1038/nature09061
  57. Ortega, F., Z. Gasparini, A. D. Buscalioni, and J. O. Calvo. 2000. A new species of Araripesuchus (Crocodylomorpha, Mesoeucrocodylia) from the Lower Cretaceous of Patagonia (Argentina). Journal of Vertebrate Paleontology 20:57–76. https://doi.org/10.1671/0272-4634(2000)020[0057:ANSOAC]2.0.CO;2
  58. Pinheiro, A. E. P., P. V. L. G. C. Pereira, R. G. de Souza, A. S. Brum, R. T. Lopes, A. S. Machado, L. P. Bergqvist, and F. M. Simbras. 2018. Reassessment of the enigmatic crocodyliform “Goniopholis” paulistanus Roxo, 1936: Historical approach, systematic, and description by new materials. PLoS ONE 13:e0199984. https://doi.org/10.1371/journal.pone.0199984
  59. Pinheiro, A. E. P., L. G. de Souza, K. L. N. Bandeira, A. S. Brum, P. V. L. G. C. Pereira, L. O. R. de Castro, R. R. C. Ramos, and F. M. Simbras. 2021. The first notosuchian crocodyliform from the Araçatuba Formation (Bauru Group, Paraná Basin), and diversification of sphagesaurians. Anais da Academia Brasileira de Ciências 93:e20201591. https://doi.org/10.1590/0001-3765202120201591
  60. Pinheiro, A. E. P., P. V. L. G. C. Pereira, F. M. Vasconcellos, A. S. Brum, L. G. de Souza, F. R. Costa, L. O. R. de Castro, K. F. da Silva, and K. L. N. Bandeira. 2023. New Itasuchidae (Sebecia, Ziphosuchia) remains and the radiation of an elusive Mesoeucrocodylia clade. Historical Biology 35:2280–2305. https://doi:10.1080/08912963.2022.2139179
  61. Pol, D. 2003. New remains of Sphagesaurus huenei (Crocodylomorpha: Mesoeucrocodylia) from the Late Cretaceous of Brazil. Journal of Vertebrate Paleontology 23(4): 817–831. https://doi.org/10.1671/A1015-7
  62. Pol, D., S. Ji, J. M. Clark, and L. M. Chiappe. 2004. Basal crocodyliforms from the Lower Cretaceous Tugulu Group (Xinjiang, China), and the phylogenetic position of Edentosuchus. Cretaceous Research 25:603–622. https://doi.org/10.1016/j.cretres.2004.05.002
  63. Pol, D. 2005. Postcranial remains of Notosuchus terrestris (Archosauria: Crocodyliformes) from the upper Cretaceous of Patagonia, Argentina. Ameghiniana 42(1):21–38.
  64. Pol, D., and S. Apesteguía. 2005. New Araripesuchus remains from the Early Late Cretaceous (Cenomanian-Turonian) of Patagonia. American Museum Novitates 3490:1–38. https://doi.org/10.1206/0003-0082(2005)490[0001:NARFTE]2.0.CO;2
  65. Pol, D., and J. E. Powell. 2011. A new sebecid mesoeucrocodylian from the Río Loro Formation (Paleocene) of north-western Argentina. Zoological Journal of the Linnean Society 163:S7–S36. https://doi.org/10.1111/j.1096-3642.2011.00714.x
  66. Pol, D., J. M. Leardi, A. Lecuona, and M. Krause. 2012. Postcranial anatomy of Sebecus icaeorhinus (Crocodyliformes, Sebecidae) from the Eocene of Patagonia. Journal of Vertebrate Paleontology 32(2):328–354. https://doi.org/10.1080/02724634.2012.646833.
  67. Pol, D., P. M. Nascimento, A. B. Carvalho, C. Riccomini, R. A. Pires-Domingues, and H. Zaher. 2014. A new notosuchian from the Late Cretaceous of Brazil and the phylogeny of advanced notosuchians. PLoS ONE 9(4):e93105. https://doi.org/10.1371/journal.pone.0093105
  68. Pol, D., and J. M. Leardi. 2015. Diversity patterns of Notosuchia (Crocodyliformes; Mesoeucrocodylia) during the Cretaceous of Gondwana. Publicación Electrónica de la Asociación Paleontológica Argentina (PE-APA) 15(1):172–186. https://doi.org/10.5710/PEAPA.10.06.2015.108
  69. Price, L. I. 1955. Novos crocodilideos dos Arenitos da Série Bauru, Cretáceo do estado de Minas Gerais. Anais da Academia Brasileira de Ciências 27:487–498.
  70. Price, L. I. 1959. Sôbre um crocodilídeo Notossúquio do Cretácico Brasileiro. Departamento Nacional da Produção Mineral, Divisão de Geologia e Mineralogia, Boletim 188:7–55.
  71. Rabi, M., and N. Sebök. 2015. A revised Eurogondwana model: Late Cretaceous notosuchian crocodyliforms and other vertebrate taxa suggest the retention of episodic faunal links between Europe and Gondwana during most of the Cretaceous. Gondwana Research 28(3): 1197–1211. https://doi.org/10.1016/j.gr.2014.09.015
  72. Ruiz, J. V., M. Bronzati, G. S. Ferreira, K. C. Martins, M. V. Queiroz, M. C. Langer, and F. Montefeltro. 2021. A new species of Caipirasuchus (Notosuchia, Sphagesauridae) from the Late Cretaceous of Brazil and the evolutionary history of Sphagesauria. Journal of Systematic Palaeontology 19:265–287. https://doi:10.1080/14772019.2021.1888815
  73. Rusconi, C. 1933. Sobre reptiles cretaceous del Uruguay (Uruguaysuchus aznarezi, n. g. n. sp.) y sus relaciones con los notosúquidos de Patagonia. Boletín Instituto de Geología y Perforaciones (Montevideo, Uruguay) 19:1–64.
  74. Sellés, A. G., A. Blanco, B. Vila, J. Marmi, F. J. López-Soriano, S. Llácer, J. Frigola, M. Canals, and A. Galobart. 2020. A small Cretaceous crocodyliform in a dinosaur nesting ground and the origin of sebecids. Scientific Reports 10:15293. https://doi.org/10.1038/s41598-020-71975-y
  75. Sereno, P. C., H. C. E. Larsson, C. A. Sidor, and B. Gado. 2001. The giant crocodyliform Sarcosuchus from the Cretaceous of Africa. Science 294:1516–1519. https://doi.org/10.1126/science.1066521
  76. Sereno, P. C. 2005. The logical basis of phylogenetic taxonomy. Systematic Biology 54(4):595–619. https://doi.org/10.1080/106351591007453
  77. Sereno, P. C., C. A. Sidor, H. C. E. Larsson, and B. Gado. 2003. A new notosuchian from the Early Cretaceous of Niger. Journal of Vertebrate Paleontology 23:477–482. https://doi.org/10.1671/0272-4634(2003)023[0477:ANNFTE]2.0.CO;2
  78. Sereno, P. C., and H. C. E. Larsson. 2009. Cretaceous crocodyliforms from the Sahara. ZooKeys 28:1–143. https://doi.org/10.3897/zookeys.28.325
  79. Sertich, J. J. W., and P. M. O’Connor. 2014. A new crocodyliform from the middle Cretaceous Galula Formation, southwestern Tanzania. Journal of Vertebrate Paleontology 34:576–596. https://doi.org/10.1080/02724634.2013.819808
  80. Simpson, G. G. 1937. New reptiles from the Eocene of South America. American Museum Novitates 927:1–3.
  81. Soto, M., D. Pol, and D. Perea. 2011. A new specimen of Uruguaysuchus aznarezi (Crocodyliformes: Notosuchia) from the Cretaceous of Uruguay and its phylogenetic relationships. Zoological Journal of the Linnean Society 163:S173–S198. https://doi.org/10.1111/j.1096-3642.2011.00717.x
  82. Steel, R. 1973. Crocodylia. Pp. 1–116 in Handbuch der Paläoherpetologie 16 (O. Kuhn, ed.). G. Fischer, Stuttgart and Portland.
  83. Turner, A. H., and J. O. Calvo. 2005. A new sebecosuchian crocodyliform from the Late Cretaceous of Patagonia. Journal of Vertebrate Paleontology 25:87–98. https://doi.org/10.1671/0272-4634(2005)025[0087:ANSCFT]2.0.CO;2
  84. Turner, A. H. 2006. Osteology and phylogeny of a new species of Araripesuchus (Crocodyliformes: Mesoeucrocodylia) from the Late Cretaceous of Madagascar. Historical Biology 18:255–369. https://doi.org/10.1080/08912960500516112
  85. Turner, A. H., and J. J. W. Sertich. 2010. Phylogenetic history of Simosuchus clarki (Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar. Journal of Vertebrate Paleontology 30(s1):177–236. https://doi.org/10.1080/02724634.2010.532348.
  86. Walker, A. D. 1968. Protosuchus, Proterochampsa, and the origin of phytosaurs and crocodiles. Geological Magazine 105:1–14. https://doi.org/10.1017/S0016756800046434
  87. Young, M. T., E. W. Wilberg, M. M. Johnson, Y. Herrera, M. B. Andrade, A. Brignon, S. Sachs, P. Abel, D. Foffa, M. S. Fernández, P. Vignaud, T. Cowgill, and S. L. Brusatte. 2024. The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha). Zoological Journal of the Linnean Society 200(2):547–617. https://doi.org/10.1093/zoolinnean/zlad165