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Issue: Vol. 19 No. 2: 21 Apr. 2026
Type: Article
Published: 2026-04-21
DOI: 10.11646/megataxa.19.2.2
Page range: 270-378
Abstract views: 92
PDF downloaded: 46

All genera of the world: Order Scorpiones (Animalia: Arthropoda: Arachnida)

Arachnology Lab and Scorpion Systematics Research Group, Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, U.S.A.
†Branchioscorpionina Neoscorpionina †Bilobosternina †Holosternina †Lobosternina †Meristosternina Orthosternina Buthida Iurida †Acanthoscorpionoidea †Branchioscorpionoidea Bothriuroidea Buthoidea Caraboctonoidea Chactoidea Chaeriloidea †Cyclophthalmoidea †Eoctonoidea †Gigantoscorpionoidea Hadruroidea †Isobuthoidea Iuroidea †Loboarchaeoctonoidea †Mesophonoidea †Palaeobuthoidea †Palaeophonoidea †Palaeoscorpioidea †Paraisobuthoidea Pseudochactoidea Scorpionoidea †Spongiophonoidea †Sucinlourencoidea Superstitionioidea Vaejovoidea †Acanthoscorpionidae Akravidae †Anthracoscorpionidae Anuroctonidae †Archaeobuthidae Belisariidae Bothriuridae †Branchioscorpionidae Buthidae Caraboctonidae †Centromachidae Chactidae Chaerilidae †Cyclophthalmidae Diplocentridae †Dolichophonidae †Eobuthidae †Eoctonidae †Eoscorpiidae Euscorpiidae †Gallioscorpionidae †Garnettiidae †Gigantoscorpionidae Hadruridae †Heloscorpionidae Hemiscorpiidae Heteroscorpionidae Hormuridae †Isobuthidae Iuridae †Kronoscorpionidae †Loboarchaeoctonidae †Mazoniidae †Mesophonidae †Microlabiidae Nullibrotheidae †Palaeoburmesebuthidae †Palaeobuthidae †Palaeoeuscorpiidae †Palaeophonidae †Palaeopisthacanthidae †Palaeoscorpiidae †Palaeotrilineatidae †Paraisobuthidae †Pareobuthidae †Praearcturidae †Proscorpiidae †Protobuthidae †Protochactidae †Protoischnuridae Pseudochactidae Rugodentidae †Scoloposcorpionidae Scorpionidae Scorpiopidae †Spongiophonidae †Stenoscorpionidae †Sucinlourencoidae Superstitioniidae †Telmatoscorpionidae Troglotayosicidae Typhlochactidae Uroctonidae Urodacidae Vaejovidae †Waterstoniidae †Willsiscorpionidae subfamilies genera synonyms type species scorpions

Abstract

The present contribution provides a consensus classification of the arachnid Order Scorpiones C.L. Koch, 1850, and updates the counts of extant and extinct genera and species through the end of 2025. Including the revisions implemented herein, there are 459 genus-group names available in Scorpiones by the end of 2025. Of these, 318 refer to currently accepted extant genera (220), subfossil genera (1) and extinct genera (97). Fifty-four genus-group names are newly synonymized, raising to 145 the number in synonymy, whereas sixteen genus-group names are revalidated and/or newly elevated to the rank of genus. Including the revisions implemented herein, Scorpiones includes 3,089 currently accepted species-group names (2,918 extant species, 1 subfossil species, and 170 extinct species) and 22 nomina dubia. Forty-seven species-group names are newly synonymized, whereas 43 species-group names are revalidated and/or newly elevated to the rank of species. 197 (62%) of the currently accepted genus-group names were described since 1950. Despite this descriptive activity, the suprageneric phylogeny and classification of Recent scorpions remains in a state of flux and largely decoupled from the phylogeny and classification of fossil forms.

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  478. Lourenço, W.R. (2000b) Un nouveau genre de scorpion malgache, maillon possible entre les Microcharmidae et les Buthidae. [A new genus of Malagasy scorpion, a possible link between the Microcharmidae and the Buthidae.] Comptes Rendus de l’Académie des Sciences, Série III Sciences de la Vie, 323 (10), 877–881. https://doi.org/10.1016/S0764-4469(00)01234-8
  479. Lourenço, W.R. (2000c) More about the Buthoidea of Madagascar, with special references to the genus Tityobuthus Pocock (Scorpiones, Buthidae). Revue suisse de Zoologie, 107 (4), 721–736. https://doi.org/10.5962/bhl.part.80145
  480. Lourenço, W.R. (2000d) Synopsis of the Colombian species of Tityus Koch (Chelicerata, Scorpiones, Buthidae), with descriptions of three new species. Journal of Natural History, 34 (3), 449–461. https://doi.org/10.1080/002229300299561
  481. Lourenço, W.R. (2000e) A propos d’une nouvelle sous-espèce géographique pour Bothriurus rochai Mello-Leitão (Scorpiones, Bothriuridae). Acta Biologica Paranaense, Curitiba, 29 (1–4), 117–125. https://doi.org/10.5380/abpr.v29i0.586
  482. Lourenço, W.R. (2001a) A remarkable scorpion fossile from the amber of Lebanon. Implication for the phylogeny of Buthoidea. Comptes Rendus de l’Academie des Sciences, Série IIA Earth and Planetary Science, 332 (10), 641–646. https://doi.org/10.1016/S1251-8050(01)01583-X
  483. Lourenço, W.R. (2001b) Un nouveau genre de Buthidae, probable vicariant géographique d’Anomalobuthus Kraepelin (Chelicerata, Scorpiones). Biogeographica (Paris), 77 (1), 15–20.
  484. Lourenço, W.R. (2001c) A propos de quelques decisions taxonomiques concernant des scorpions des families Buthidae, Chaerilidae et Scorpiopidae. Biogeographica (Paris), 77 (4), 173–175.
  485. Lourenço, W.R. (2001d) Un nouveau genre et une nouvelle espèce de scorpion d’Algérie, avec des considérations taxonomiques sur le genre Lissothus Vachon, 1948 (Scorpiones, Buthidae). Zoosystema, 23 (1), 51–57.
  486. Lourenço, W.R. (2001e) Nouvelles considérations sur la phylogénie et la biogéographie des scorpions Ischnuridae de Madagascar. Biogeographica (Paris), 77 (2), 83–96.
  487. Lourenço, W.R. (2001f) Taxonomic considerations on the genera Butheolus Simon, Nanobuthus Pocock and Neobuthus Hirst (Scorpiones, Buthidae) with the description of a new species of Neobuthus from Ethiopia. In: Prakash I. (Ed.), Ecology of desert environments. A Festschrift for Prof. J.L. Cloudsley-Thompson on his 80th birthday. Scientific Publishers, Jodhpur, India, pp. 171–183.
  488. Lourenço, W.R. (2002a) The first scorpion fossil from the Cretaceous amber of Myanmar (Burma). New implications for the phylogeny of Buthoidea. Comptes Rendus Palevol, 1 (2), 97–101. https://doi.org/10.1016/S1631-0683(02)00017-9
  489. Lourenço, W.R. (2002b) Nouvelles additions à la faune de scorpions Néotropicaux (Arachnida). Revue suisse de Zoologie, 109 (1), 127–141. https://doi.org/10.5962/bhl.part.79582
  490. Lourenço, W.R. (2002c) Nouvelles considérations sur la systématique et la biogéographie du genre Butheoloides Hirst (Scorpiones, Buthidae) avec description d’un nouveau sous-genre et de deux nouvelles espèces. Revue suisse de Zoologie, 109 (4), 725–733.
  491. Lourenço, W.R. (2003a) The first scorpion fossil from the Cretaceous amber of France. New implications for the phylogeny of Chactoidea. Comptes Rendus Palevol, 2 (3), 213–219. https://doi.org/10.1016/S1631-0683(03)00042-3
  492. Lourenço, W.R. (2003b) Humicolous buthoid scorpions: A new genus and species from French Guiana. Comptes Rendus Biologies, 326 (12), 1149–1155. https://doi.org/10.1016/j.crvi.2003.09.003
  493. Lourenço, W.R. (2003c) About two species of liochelid scorpions collected in western Africa (Scorpiones, Liochelidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (168), 137–148.
  494. Lourenço, W.R. (2004) Humicolous microcharmid scorpions: A new genus and species from Madagascar. Comptes Rendus Biologies, 327 (1), 77–83. https://doi.org/10.1016/j.crvi.2003.12.003
  495. Lourenço, W.R. (2005a) A new genus and species of scorpion from Afghanistan (Scorpiones, Buthidae). Bonner zoologische Beiträge, 53 (1/2), 111–114.
  496. Lourenço, W.R. (2005b) Description of three new species of scorpion from Sudan (Scorpiones, Buthidae). Boletín de la Sociedad Entomológica Aragonesa, 36, 21–28.
  497. Lourenço, W.R. (2005c) Scorpion diversity and endemism in the Rio Negro region of Brazilian Amazonia, with the description of two new species of Tityus C.L. Koch (Scorpiones, Buthidae). Amazoniana, 18 (3–4), 203–213.
  498. Lourenço, W.R. (2005d) A new species of Tityus Koch, 1836 (Scorpiones, Buthidae) from Ecuador. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (171), 221–227.
  499. Lourenço, W.R. (2006) Une nouvelle proposition de découpage sous-générique du genre Tityus C. L. Koch, 1836 (Scorpiones, Buthidae). Boletín de la Sociedad Entomológica Aragonesa, 39, 55–67.
  500. Lourenço, W.R. (2007a) A new species of Tityus C.L. Koch, 1836 from Ecuador: The first element of the ‘Tityus androcottoides’ subgroup for this country. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (176), 375–385.
  501. Lourenço, W.R. (2007b) First record of the family Pseudochactidae Gromov (Chelicerata, Scorpiones) from Laos and new biogeographic evidence of a Pangaean palaeodistribution. Comptes Rendus Biologies, 330 (10), 770–777. https://doi.org/10.1016/j.crvi.2007.07.006
  502. Lourenço, W.R. (2007c) Further additions to the scorpion fauna of the Guayana region of South America. Revue suisse de Zoologie, 114 (3), 513–519. https://doi.org/10.5962/bhl.part.80402
  503. Lourenço, W.R. (2008a) The geographic pattern of distribution of the genus Rhopalurus Thorell, 1876 in the Guayana-Amazon region (Scorpiones: Buthidae). Euscorpius, 73, 1–14. https://doi.org/10.18590/euscorpius.2008.vol2008.iss73.1
  504. Lourenço, W.R. (2008b) Description of Tityus (Atreus) neblina sp. n. (Scorpiones, Buthidae), from the ‘Parque Nacional do Pico da Neblina’, in Brazil/Venezuela, with comments on some related species. Boletín de la Sociedad Entomológica Aragonesa, 43, 75–79.
  505. Lourenço, W.R. (2009) Eyeless forest litter scorpions; a new species from the island of Halmahera (Moluccas), Indonesia (Scorpiones, Chaerilidae). Boletín de la Sociedad Entomológica Aragonesa, 44, 93–97.
  506. Lourenço, W.R. (2012a) About the scorpion fossils from the Cretaceous amber of Myanmar (Burma) with the descriptions of a new family, genus and species. Acta Biologica Paranaense, Curitiba, 41 (3–4), 75–87.
  507. Lourenço, W.R. (2012b) The genus Vietbocap Lourenço and Pham, 2010 (Scorpiones: Pseudochactidae); proposition of a new subfamily and description of a new species from Laos. Comptes Rendus Biologies, 335 (3), 232–237. https://doi.org/10.1016/j.crvi.2012.02.001
  508. Lourenço, W.R. (2012c) Further considerations on Tityus (Archaeotityus) clathratus C. L. Koch, 1844 and description of two associated new species (Scorpiones, Buthidae). Boletín de la Sociedad Entomológica Aragonesa, 50, 277–283.
  509. Lourenço, W.R. (2013) A new subgenus and species of Alloscorpiops Vachon, 1980 from Laos (Scorpiones, Euscorpiidae, Scorpiopinae); implications for the taxonomy of the group. Comptes Rendus Biologies, 336 (1), 51–55. https://doi.org/10.1016/j.crvi.2013.01.001
  510. Lourenço, W.R. (2014a) A new species of scorpion from Chiapas amber, Mexico (Scorpiones: Buthidae). Revista Ibérica de Aracnología, 24, 59–63.
  511. Lourenço, W.R. (2014b) The genus Broteochactas Pocock, 1893 in Brazilian Amazonia, with a description of a new species from the State of Amazonas (Scorpiones: Chactidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 17 (192), 153–159.
  512. Lourenço, W.R. (2014c) Micro-endemic and vicariant populations of Opisthacanthus Peters, 1861 (Scorpiones: Hormuridae) in Madagascar, with descriptions of two new species. Arthropoda Selecta, 23 (4), 383–389. https://doi.org/10.15298/arthsel.23.4.07
  513. Lourenço, W.R. (2015a) A new subfamily, genus and species of fossil scorpions from Cretaceous Burmese amber (Scorpiones, Palaeoeuscorpiidae). In: Wunderlich, J. (Ed.), Mesozoic spiders (Araneae): Ancient spider faunas and spider evolution. Beiträge zur Araneologie, 9, 457–464.
  514. Lourenço, W.R. (2015b) Clarification of the familiar status of the genus Palaeoburmesebuthus Lourenço, 2002 from Cretaceous Burmese amber (Scorpiones: Archaeobuthidae: Palaeoburmesebuthinae). In: Wunderlich, J. (Ed.), Mesozoic spiders (Araneae): Ancient spider faunas and spider evolution. Beiträge zur Araneologie, 9, 465–475.
  515. Lourenço, W.R. (2015c) New considerations on the enigmatic genus Vachonus Tikader et Bastawade, 1983, with the description of a new species from India (Scorpiones: Buthidae). Arachnida – Rivista Aracnologica Italiana, 3, 2–9.
  516. Lourenço, W.R. (2016a) A complement of information about the status of the genus Sabinebuthus Lourenço, 2001 (Scorpiones: Buthidae). Arachnida – Rivista Aracnologica Italiana, 10, 18–25.
  517. Lourenço, W.R. (2016b) Scorpions from the Mitaraka Massif in French Guiana: Description of one new genus and species (Scorpiones: Chactidae). Comptes Rendus Biologies, 339 (3–4), 141–146. https://doi.org/10.1016/j.crvi.2016.02.003
  518. Lourenço, W.R. (2016c) A new genus and three new species of scorpions from Cretaceous Burmese amber (Scorpiones, Chaerilobuthidae, Palaeoeuscorpiidae). Arthropoda Selecta, 25 (1), 67–74. https://doi.org/10.15298/arthsel.25.1.06
  519. Lourenço, W.R. (2017a) One more new species of Broteochactas Pocock, 1893 (Scorpiones: Chactidae) from Brazilian Amazonia. Revista Ibérica de Aracnología, 30, 11–14.
  520. Lourenço, W.R. (2017b) Scorpions from Brazilian Amazonia, with a description of two new species from ‘Serra da Mocidade’ National Park in the State of Roraima (Scorpiones: Buthidae, Chactidae). Arachnida – Rivista Aracnologica Italiana, 12, 2–17.
  521. Lourenço, W.R. (2017c) A new genus and species of scorpion from Burma [Myanmar] (Scorpiones: Scorpiopidae): Implications for the taxonomy of the family. Comptes Rendus Biologies, 340 (6–7), 349–357. https://doi.org/10.1016/j.crvi.2017.05.003
  522. Lourenço, W.R. (2018) A new remarkable scorpion genus and species from Cretaceous Burmese amber (Scorpiones: Protoischnuridae). Arachnida – Rivista Aracnologica Italiana, 18, 2–14.
  523. Lourenço, W.R. (2019) New insights on the scorpion species of the Tityus trivittatus group of subgenus Tityus C. L. Koch, 1836 (Scorpiones: Buthidae). Revista Ibérica de Aracnología, 34, 119–125.
  524. Lourenço, W.R. (2020) A possible relict population of Mesobuthus (Aegaeobuthus?) nigrocinctus (Ehrenberg, 1828) in the Bishri Mountains of Syria (Scorpiones: Buthidae). Serket, 17 (2), 77–86.
  525. Lourenço, W.R. (2021a) A further new species for the Malagasy genus Pseudouroplectes Lourenço, 1995 (Scorpiones: Buthidae). Faunitaxys, 9 (41), 1–7. https://doi.org/10.57800/faunitaxys-9(41)
  526. Lourenço, W.R. (2021b) The genus Microananteris Lourenço, 2003 in French Guiana (Scorpiones: Buthidae). Zoosystema, 43 (20), 377–386. https://doi.org/10.5252/zoosystema2021v43a20
  527. Lourenço, W.R. (2023) Nouvelles considérations sur Isometrus (Reddyanus) heimi Vachon, 1976 et description d’une deuxième espèce d’Isometrus pour la Nouvelle-Calédonie (Scorpiones: Buthidae). Revue Arachnologique, 2 (10), 30–36.
  528. Lourenço, W.R. & Araújo, J. de S. (2004) Nouvelles considérations sur le genre Auyantepuia González-Sponga, 1978 (Scorpiones, Chactidae) et description d’une nouvelle espèce pour la région de Manaus, en Amazonie brésilienne. Acta Biologica Paranaense, 33 (1/4), 1–11. https://doi.org/10.5380/abpr.v33i0.623
  529. Lourenço, W.R. & Beigel, A. (2011) A new scorpion fossil from the Cretaceous amber of Myanmar (Burma). New phylogenetic implications. Comptes Rendus Palevol, 10 (8), 635–639. https://doi.org/10.1016/j.crpv.2011.08.001
  530. Lourenço, W.R. & Beigel, A. (2015) A new genus and species of Palaeoburmesebuthinae Lourenço, 2015 (Scorpiones, Archaeobuthidae) from Cretaceous amber of Myanmar. In: Wunderlich, J. (Ed.), Mesozoic spiders (Araneae): Ancient spider faunas and spider evolution. Beiträge zur Araneologie, 9, 476–480.
  531. Lourenço, W.R. & Bruehmueller Ramos, E.C. (2004) New considerations on the status of Tityus magnimanus Pocock, 1897 (Scorpiones, Buthidae) and description of a new species of Tityus from the State of Roraima, Brazil. Revista Ibérica de Aracnología, 10, 285–291.
  532. Lourenço, W.R. & Chevalier, J. (2022) A further new species of the genus Microananteris Lourenço, 2003, from French Guiana (Scorpiones, Buthidae). Bulletin de la Société entomologique de France, 127 (1), 91–99. https://doi.org/10.32475/bsef_2228
  533. Lourenço, W.R. & Cloudsley-Thompson, J.L. (1995) Stridulatory apparatus and the evolutionary significance of sound production in Rhopalurus species (Scorpiones: Buthidae). Journal of Arid Environments, 31 (4), 423–429. https://doi.org/10.1016/S0140-1963(05)80125-9
  534. Lourenço, W.R. & Cloudsley-Thompson, J.L. (1996a) The evolutionary significance of colour, colour patterns and fluorescence in scorpions. Revue suisse de Zoologie, hors série, 2, 449–458.
  535. Lourenço, W.R. & Cloudsley-Thompson, J.L. (1996b) Recognition and distribution of the scorpions of the genus Pandinus Thorell, 1876 accorded protection by the Washington convention. Biogeographica (Paris), 72, 133–143.
  536. Lourenço, W.R. & Duhem, B. (2009) Saharo-Sindian buthid scorpions; description of two new genera and species from Occidental Sahara and Afghanistan. ZooKeys, 14, 37–54. https://doi.org/10.3897/zookeys.14.212
  537. Lourenço, W.R. & Duhem, B. (2010) A new species of Auyantepuia Gonzáles-Sponga, 1978 (Scorpiones, Chactidae) from Suriname. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 15 (182), 137–146.
  538. Lourenço, W.R. & Flórez, E. (1990) Scorpions (Chelicerata) from Colombia. III. The scorpio-fauna of the Pacific Region (Choco), with some biogeographic considerations. Amazoniana, 11 (2), 119–134.
  539. Lourenço, W.R. & Flórez D., E. (2018) Further clarifications on some species of the genus Tityus, subgenus Atreus (Scorpiones: Buthidae), from Colombia and Ecuador, with the description of a new species. Revista Ibérica de Aracnología, 33, 63–72.
  540. Lourenço, W.R. & Gall, J.C. (2004) Fossil scorpions from the Buntsandstein (Early Triassic) of France. Comptes Rendus Palevol, 3 (5), 369–378. https://doi.org/10.1016/j.crpv.2004.06.006
  541. Lourenço, W.R. & Goodman, S.M. (1999) Taxonomic and ecological observations on the scorpions collected in the Réserve Naturelle Intégrale d’Andohahela, Madagascar. In: Goodman, S.M. (Ed.), A floral and faunal inventory of the Réserve Naturelle Intégrale d’Andohahela, Madagascar: With reference to elevational variation. Fieldiana Zoology, new series, 94, 149–153.
  542. Lourenço, W.R. & Goodman, S.M. (2006) A reappraisal of the geographical distribution of the genus Opisthacanthus Peters, 1861 (Scorpiones: Liochelidae) in Madagascar, including the description of four new species. Boletín de la Sociedad Entomológica Aragonesa, 38, 11–23.
  543. Lourenço, W.R. & Goodman, S.M. (2008) Scorpions of the Réserve spéciale d’Ankarana, Madagascar, with particular reference to cave-dwelling animals and the description of two new species (Arachnida, Scorpiones). Zoosystema, 30 (3), 665–679.
  544. Lourenco, W.R. & Košulič, O. (2018) A new remarkable species of Alloscorpiops Vachon, 1980 from Myanmar (Burma) (Scorpiones, Scorpiopidae). ZooKeys, 775, 47–58. https://doi.org/10.3897/zookeys.775.24248
  545. Lourenço, W.R. & Leguin, E.-A. (2025) New comments on the scorpion diversity of Ecuador and description of a new species of Tityus C. L. Koch, 1836 (Scorpiones: Buthidae). Revista Ibérica de Aracnología, 46, 27–32.
  546. Lourenço, W.R. & Monod, L. (2000) Description of a new genus and species of scorpion (Bothriuridae) from Brazil. Ekologia (Bratislava), 19 (Suppl. 3), 145–152.
  547. Lourenço, W.R. & Otero Patiño, R. (1998) Tityus antioquensis sp. n., a new species of scorpion from the Department Antioquia, Central Cordillera of Colombia (Scorpiones, Buthidae), with a checklist and key for the Colombian species of the genus. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 12 (158), 297–307.
  548. Lourenço, W.R. & Pézier, A. (2002) Addition to the scorpion fauna of the Manaus region (Brazil), with a description of two new species of Tityus from the canopy. Amazoniana, 17 (1), 177–186.
  549. Lourenço, W.R. & Pham, D.-S. (2010) A remarkable new cave scorpion of the family Pseudochactidae Gromov (Chelicerata, Scorpiones) from Vietnam. ZooKeys, 71, 1–13. https://doi.org/10.3897/zookeys.71.786
  550. Lourenço, W.R. & Pham, D.-S. (2015a) An interesting new subgenus of Scorpiops Peters, 1861 from north Vietnam (Scorpiones: Euscorpiidae: Scorpiopinae). Comptes Rendus Biologies, 338 (3), 212–217. https://doi.org/10.1016/j.crvi.2015.01.003
  551. Lourenço, W.R. & Pham, D.-S. (2015b) A new remarkable species of Alloscorpiops Vachon, 1980 from a cave in Vietnam (Scorpiones, Euscorpiidae, Scorpiopinae). ZooKeys, 500, 73–82. https://doi.org/10.3897/zookeys.500.9421
  552. Lourenço, W.R. & Qi, J.-X. (2006) Mountain scorpions: A new genus and species from Tibet (China). Comptes Rendus Biologies, 329 (4), 289–295. https://doi.org/10.1016/j.crvi.2006.02.005
  553. Lourenço, W.R. & Qi, J.-X. (2007a) Additions à la faune des scorpions de l’Etat du Amapá, Brésil (Chelicerata, Scorpiones). Revue suisse de Zoologie, 114 (1), 3–12. https://doi.org/10.5962/bhl.part.80384
  554. Lourenço, W.R. & Qi, J.-X. (2007b) Scorpions from the rainforest canopy of New Guinea and description of a new subspecies of Lychas C. L. Koch, 1845 (Scorpiones: Buthidae). Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, Entomologie, 77, 157–161.
  555. Lourenço, W.R. & Rossi, A. (2019) Further clarifications on species of Tityus C. L. Koch, 1836, subgenus Atreus Gervais, 1843 (Scorpiones: Buthidae), from Amazonia, with the description of a new species. Arachnida – Rivista Aracnologica Italiana, 21, 1–24.
  556. Lourenço, W.R. & Vachon, M. (1981) Compléments à la déscription d’Acanthothraustes brasiliensis (Mello-Leitão 1931) (= Teuthraustes brasilianus Mello-Leitão 1931), synonyme d’Euscorpius flavicaudis (Geer 1778) (Scorpiones, Chactidae). Journal of Arachnology, 9 (2), 223–228.
  557. Lourenço, W.R. & Vachon, M. (1995) Un nouveau genre et deux nouvelles espèces de scorpions Buthidae d’Iran. Bulletin du Musèum National d’Histoire Naturelle, Paris (Zoologie, Biologie et Écologie Animale), (4), 17, A, (3–4), 297–305. https://doi.org/10.5962/p.290355
  558. Lourenço, W.R. & Vachon, M. (1997) Un nouveau genre et quatre nouvelles espèces de scorpions (Buthidae) du Moyen-Orient. Zoosystema, 19 (2–3), 327–336. https://doi.org/10.5962/p.296162
  559. Lourenço, W.R. & Velten, J. (2016) A new genus and species of fossil scorpion from Burmese Cretaceous amber (Scorpiones: Buthoidea: Buthidae). Arachnida – Rivista Aracnologica Italiana, 10, 2–9.
  560. Lourenço, W.R. & Velten, J. (2017) One more new genus and species of fossil scorpion from Burmese Cretaceous amber belonging to the Palaeoburmesebuthidae (Scorpiones). Arachnida – Rivista Aracnologica Italiana, 13, 2–10.
  561. Lourenço, W.R. & Velten, J. (2020) A new contribution to the knowledge of Cretaceous Burmese amber scorpions with the descriptions of two new genera and two new species (Scorpiones: Chaerilobuthidae; Palaeoeuscorpiidae: Archaeoscorpiopinae). Arachnida – Rivista Aracnologica Italiana, 28, 34–50.
  562. Lourenço, W.R. & Velten, J. (2021) One more new genus and species of scorpion from Early Cretaceous Burmese amber (Scorpiones: Protoischnuridae). Faunitaxys, 9 (14), 1–5. https://doi.org/10.57800/faunitaxys-9(14)
  563. Lourenço, W.R. & Velten, J. (2022) Further insights on Cretaceous Burmite scorpions with the descriptions of a new genus and species (Scorpiones: Buthoidea: Buthidae). Faunitaxys, 10 (35), 1–5. https://doi.org/10.57800/faunitaxys-10(35)
  564. Lourenço, W.R. & Velten, J. (2024a) An exceptional new genus of fossil scorpion from Burmese Cretaceous amber belonging to the family Palaeoburmesebuthidae Lourenço, 2015. Faunitaxys, 12 (13), 1–7. https://doi.org/10.57800/faunitaxys-12(13)
  565. Lourenço, W.R. & Velten, J. (2024b) New insights on the diversity of the family Chaerilobuthidae Lourenço and Beigel, 2011 with the description of one new genus and species (Scorpiones). Faunitaxys, 12 (46), 1–6. https://doi.org/10.57800/faunitaxys-12(46)
  566. Lourenço, W.R. & Velten, J. (2025a) Description of a second species for the genus Serratochaerilobuthus Lourenço, 2024 (Scorpiones: Chaerilobuthidae). Faunitaxys, 13 (10), 1–6. https://doi.org/10.57800/faunitaxys-13(10)
  567. Lourenço, W.R. & Velten, J. (2025b) One more remarkable new genus and species of fossil scorpion from Burmese Cretaceous amber associated to the extant family Buthidae (Scorpiones: Buthoidea). Faunitaxys, 13 (35), 1–6. https://doi.org/10.57800/faunitaxys-13(35)
  568. Lourenço, W.R. & Von Eickstedt, V.R.D. (1987) Contribução ao conhecimento taxinômico dos escorpiões associés au groupe Tityus melanostictus Pocock, 1893 (Scorpiones, Buthidae). Memórias do Instituto de Butantan, 49 (3), 87–95.
  569. Lourenço, W.R. & Weitschat, W. (1996) More than 120 years after its description, the enigmatic status of the genus of the Baltic amber scorpion “Tityus eogenus” Menge, 1869 can finally be clarified. Mitteilungen aus dem Geologisch-Paläontologischen Institut der Universität Hamburg, 79, 189–193.
  570. Lourenço, W.R. & Weitschat, W. (2000) New fossil scorpions from the Baltic amber. Implications for Cenozoïc biodiversity. Mitteilungen aus dem Geologisch-Paläontologischen Institut der Universität Hamburg, 84, 247–260.
  571. Lourenço, W.R. & Weitschat, W. (2001) Description of another fossil scorpion from Baltic amber, with considerations on the evolutionary levels of Cenozoic Buthoidea. Mitteilungen aus dem Geologisch-Paläontologischen Institut der Universität Hamburg, 85, 277–283.
  572. Lourenço, W.R. & Weitschat, W. (2005) A new genus and species of a fossil scorpion from a different kind of Baltic amber (Scorpiones, Buthidae). Mitteilungen aus dem Geologisch-Paläontologischen Institut der Universität Hamburg, 89, 183–188.
  573. Lourenço, W.R. & Wilmé, L. (2019) Scorpions from the Parc National du Tsingy de Namoroka, Madagascar and description of a new species of Opisthacanthus Peters, 1861 (Scorpiones: Hormuridae). Revista Ibérica de Aracnología, 34, 13–20.
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