A revision of the former family Pseudopomyzidae (Diptera, Acalyptratae, Cypselosomatidae)

TIFFANY YAU — 2025-12-01

The former family Pseudopomyzidae is revised. Morphological characters and multiple genes are used to assess relationships among the families of Nerioidea, concluding that the family Pseudopomyzidae as currently defined is paraphyletic with respect to Cypselosomatidae Hendel; the family Cypselosomatidae is therefore redefined to include the Pseudopomyzidae. The subgeneric classification of the large genus Pseudopomyza Strobl is updated by synonymizing the subgenera Macalpinella Papp and Rhinopomyzella Hennig with Pseudopomyza s.s., and reinstating the subgenus Heluscolia (Harrison), resulting in four valid subgenera: Apops McAlpine, Dete McAlpine, Heluscolia and Pseudopomyza s.s. The subgenus Apops is composed of two previously included species (P. flavitarsis (Harrison) (New Zealand) and P. arenae McAlpine (Australia, Tasmania)) and three new species (P. chilensis Yau and Marshall (Chile), P. medianentis Yau and Marshall (Australia, Western Australia) and P. nigritarsis Yau and Marshall (New Zealand)). The subgenus Dete includes P. collessi McAlpine (Australia). The subgenus Heluscolia includes P. antipoda (Harrison) and P. brevis (Harrison), both from the New Zealand Subantarctic Islands. The subgenus Pseudopomyza includes P. aristata (Harrison) (New Zealand), P. atrimana (Meigen) (Palaearctic), P. brevicaudata (Harrison) (New Zealand), P. brevifacies (Papp) (Oriental and Palaearctic) and the P. nigrimana species group (formerly treated as the subgenus Rhinopomyzella). The P. nigrimana species group, shown here to be the sister group to P. atrimana, includes P. albimana (Hennig), P. nigrimana (Hennig) and fifteen new species (P. adunca Yau and Marshall, P. ampliata Yau and Marshall, P. angustifrons Yau and Marshall, P. binaevia Yau and Marshall, P. brunneicacumen Yau and Marshall, P. cordata Yau and Marshall, P. cyathiformis Yau and Marshall, P. flavicacumen Yau and Marshall, P. gambiformis Yau and Marshall, P. inflexa Yau and Marshall, P. machaera Yau and Marshall, P. parabinaevia Yau and Marshall, P. proboscis Yau and Marshall, P. prolata Yau and Marshall, P. simulatrix Yau and Marshall).

**Taxonomic revision of the spider genus Azilia Keyserling, 1881 (Araneae: Tetragnathidae)**

ANTONIO D. BRESCOVIT — 2025-12-01

The species of the spider genus Azilia Keyserling, 1881 from the Americas are revised. Cardimia Mello-Leitão, 1940 is removed from the synonymy with Azilia and is newly synonymized with Diphya Nicolet, 1849, with its type species, Cardimia eximia Mello-Leitão, 1940, considered synonymous of Diphya bicolor Vellard, 1926. The genus type species, Azilia formosa Keyserling 1881 described from Peru is redescribed. The type material is considered lost, but here a neotype is designed based on specimens next to the type locality in Peru. Azilia boudeti Simon, 1895, Azilia marmorata Mello-Leitão, 1948, Arochoides integrans Mello-Leitão, 1935 and Cyrtophora vachoni Caporiacco, 1954 had their types examined and are synonymized with Azilia histrio Simon, 1895. The males of Azilia formosa and A. histrio are described for the first time. Azilia affinis O. Pickard-Cambridge, 1893, A. guatemalensis O. Pickard-Cambridge, 1889, A. rojasi Simon, 1895 and A. montana Bryant, 1940 are redescribed and illustrated. Finally, eight new species are described based on males and females, including six from Brazil: Azilia arrebitada sp. nov.; A. catita sp. nov.; A. fabulosa sp. nov.; A. guapa sp. nov.; A. joinha sp. nov.; A. supimpa sp. nov. and two from Mexico: Azilia esplendida sp. nov., and A. hermosa sp. nov.

Megataxa

A revision of the former family Pseudopomyzidae (Diptera, Acalyptratae, Cypselosomatidae)

Taxonomic revision of the spider genus Azilia Keyserling, 1881 (Araneae: Tetragnathidae)

**Taxonomic revision of the spider genus Azilia Keyserling, 1881 (Araneae: Tetragnathidae)**