Palaeoentomology <p><strong>Palaeoentomology </strong>is the official journal of the <a href="">International Palaeoentomological Society</a> (IPS). It is an international peer-reviewed scientific journal, which publishes high quality, original research contributions as well as review papers. Papers are published in English and they cover a wide spectrum of topics in palaeoentomology, fossil terrestrial arthropods and amber research, i.e. systematic palaeontology, morphology, diversity, palaeogeography, palaeoecology, palaeobehavior, evolutionary and phylogenetic studies on fossil insects and terrestrial arthropods, biostratigraphy, taphonomy, and amber (deposits, inclusions, geochemistry, curation). Descriptions of new methods (analytical, instrumental or numerical) should be relevant to the broad scope of the journal.</p> <p> </p> <p>Palaeoentomology is the flag journal of IPS, who is responsible for the editing of this journal. For more info about IPS, please contact Prof. Dr. Hab. Dany Azar, Lebanese University, Lebanon.</p> Magnolia press en-US Palaeoentomology 2624-2826 <span lang="EN-GB">Authors need to complete and return an </span><span lang="EN-GB"><a href="/phytotaxa/images/copyright.rtf">Assignment of Copyright</a> </span><span lang="EN-GB">form when a paper is accepted for publication. Authors from institutions that do not allow transfer of copyrights to publishers (e.g. government institutions such as USDA, CSIRO) should attach a copyright waiver or similar document.</span> <p><strong>A new genus of Ipsviciidae (Hemiptera, Cicadomorpha) with a tegminal strigil from the Triassic of Madygen</strong></p> <p>The family Ipsviciidae, originally described from the Upper Triassic of Australia and considered an offshoot of the Scytinopteridae (Tillyard, 1919), was later variously assigned to Heteroptera, Coleorrhyncha, Fulgoroidea, or Cicadomorpha (Cercopoidea or its own superfamily) by different authors (see Lambkin, 2020). Shcherbakov (1984) placed Ipsviciidae in the Scytinopteroidea and demonstrated that this superfamily is ancestral to Heteroptera (Shcherbakov, 1996). The family comprises several genera known from the Triassic and Lower Jurassic of Australia and Eurasia (Lambkin, 2020). A peculiar monotypic genus of Ipsviciidae with a strigil (stridulatory area) on the underside of the tegmen is described below from the Middle to Upper Triassic (Ladinian–Carnian) of the Madygen Lagerstätte, Central Asia. Such a strigil indicates that the new species possessed a stridulatory device of the forewing-hindleg type, similar to those occurring in the extinct Dysmorphoptilidae (Evans, 1961) and some extant true bugs (see Discussion). Dysmorphoptilids and the new ipsviciid genus may have used these devices to produce alarm signals.</p> DMITRY E. SHCHERBAKOV Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 434–438 434–438 10.11646/palaeoentomology.5.5.3 <p><strong>A proctotrupid wasp in Lebanese Lower Cretaceous amber (Hymenoptera: Proctotrupidae)</strong></p> <p>The first fossil proctotrupid wasp in Lower Cretaceous Lebanese amber is described and figured. <em>Astarteserphus grimaldii</em> <strong>gen. et sp. nov.</strong> is distinguished from all other living and fossil Proctotrupidae and placed into its own subfamily, Astarteserphinae <strong>subfam. nov.</strong> A key is presented to the living and fossil subfamilies and tribes of Proctotrupidae, and a brief discussion is provided on the unique features and diminutive size of the fossil.</p> MICHAEL S. ENGEL HOLLISTER W. HERHOLD PHILLIP BARDEN Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 439–444 439–444 10.11646/palaeoentomology.5.5.4 <p><strong>The first Proctotrupidae (Hymenoptera) in Burmese amber, with description of a new genus and species</strong></p> <p>Cresogmus grimaldii Rasnitsyn &amp; Kolyada gen. et sp. nov. is described from the mid-Cretaceous Burmese amber in the tribe Disogmini (subfamily Proctotrupinae) as the first representative of the family Proctotrupidae encased in Burmese amber. This new genus is diagnosed by the occipital carina extending to the lower half of the head, the pronotum lacking an epomia, the pterostigma short, the petiole open but very short, the ovipositor wide and bent throughout, with its apex widely rounded. Diagnostic characters of Disogmini are revised.</p> ALEXANDR P. RASNITSYN VIKTOR A. KOLYADA DMITRY D. VORONTSOV CHRISTOPH ÖHM-KÜHNLE Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 445–451 445–451 10.11646/palaeoentomology.5.5.5 <p><strong>Transitional morphology and Afrotropical affinity of a bythinoplectine rove beetle from the early Eocene of India (Coleoptera: Staphylinidae: Pselaphinae)</strong></p> <p>Recently discovered Ypresian-age amber from Cambay, India, reveals an ancient arthropod assemblage on the Indian subcontinent during its collision with Asia. Despite the tectonic history of India, limited connections have been found between the Cambay palaeofauna and present-day Madagascan and mainland African faunas. Here, I describe a new fossil pselaphine rove beetle (Staphylinidae: Pselaphinae) recovered from the Cambay deposit that shows closest apparent phylogenetic affinity to modern Afrotropical genera. <em>Yprezethinus grimaldii</em> <strong>gen. et sp. nov.</strong> is placed in Bythinoplectini, subtribe Bythinoplectina. Based on antennal and maxillary palp morphology, <em>Yprezethinus </em>is a putative transitional stem lineage of the <em>Zethinus</em>-group of genera—an extant clade distributed across equatorial African rainforests. Although <em>Yprezethinus</em> shares with this clade the derived feature of ovoid antennal clubs formed by tight appression of the apical two segments, it differs from its putative extant relatives in its possession of the plesiomorphic complement of 11 antennomeres, without any fusions of segments. The fossil taxon signifies a biotic link between early Eocene India and continental Africa, and marks the Cenozoic emergence of a tropical leaf litter arthropod fauna approaching that of contemporary, ant-dominated rainforests.</p> JOSEPH PARKER Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 452–460 452–460 10.11646/palaeoentomology.5.5.6 <p><strong>A review of the Cretaceous genus <em>Eltxo</em> (Diptera: Cecidomyiidae) with description of the new species <em>Eltxo</em> grimaldii from El Soplao amber</strong></p> <p>A new species, <em>Eltxo grimaldii</em><strong> sp. nov.</strong>, is described from Spanish Lower Cretaceous (middle Albian) amber from El Soplao, based on a single female. The new species is compared with the other only known species of the genus, <em>Eltxo cretaceus </em>Arillo &amp; Nel, 2000, based on a single male specimen also found in Spanish amber, but slightly younger (Peñacerrada I amber; upper Albian). The holotype of <em>E. cretaceus</em> is reviewed and its description corrected and expanded, providing the first micrographs of its anatomical features. The holotype of the new fossil species is the only female specimen known of the cecidomyiid tribe Amediini Jaschhof, 2021, a tribe recently described after changes of taxonomic attribution of the genus <em>Eltxo</em> during the last 20 years.</p> ENRIQUE PEÑALVER ANTONIO ARILLO ANDRÉ NEL Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 461–467 461–467 10.11646/palaeoentomology.5.5.7 <p><strong>Palleptoceridae fam. nov., an extinct leptoceroid family in mid-Cretaceous Burmese amber (Insecta, Trichoptera)</strong></p> <p>The family Palleptoceridae <strong>fam. nov.</strong> is proposed as a new extinct family of the superfamily Leptoceroidea. It is characterized by the absence of ocelli, the presence of five-segmented maxillary palps in both sexes, antennae longer than the forewings, and the tibial spur in the form of 2/4/4. The extinct Palleptoceridae is closely related to the leptoceroid family Leptoceridae, but the formula for the adult tibial spur is reduced from 2/4/4 to 2/2/4 (Morse, 1981). The <em>Palleptocerus grimaldii</em> <strong>sp. nov.</strong> from the mid-Cretaceous Burmese amber is furthermore characterized by the sexual dimorphic wing venation. In the forewings, the apical forks I and V are present in the male and forks I, III, and V in the female; in the hind wings of both sexes, exclusively the apical fork V is present. The Leptoceroidea originated and evolved in Gondwana. <em>Palleptocerus grimaldii</em> <strong>sp. nov.</strong> is interpreted as a relict species of the Gondwanan Leptoceroidea in Burmese amber.</p> WILFRIED WICHARD PATRICK MÜLLER Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 468–474 468–474 10.11646/palaeoentomology.5.5.8 <p><strong>Dolichopodidae (Diptera) from the Eocene amber deposits of Cambay and Kutch Basins, India</strong></p> <p>The family Dolichopodidae (Diptera) from the Eocene amber of the Cambay and Kutch, India is reviewed. Based on a sample of 38 inclusions, the fauna preserved in amber reveals that the family was well-differentiated by early the Eocene. <em>Palaeomedeterus cambayensis</em> Bickel <strong>sp. nov.</strong> is described and provisionally assigned to the Peloropeodinae, and is congeneric with European Baltic amber species, suggesting the genus was widespread for a considerable time period in both Europe and India. Two small-sized new monotypic genera are also described, <em>Gujaratmyia rotunda</em> Bickel <strong>gen. et sp. nov.</strong> and <em>Rajpardia grimaldii </em>Bickel <strong>gen. et sp. nov.</strong> Also included are undescribed members of the dolichopodid subfamilies Diaphorinae and Medeterinae, and unplaced species. The Baltic amber species <em>Prohercostomus noxialis</em> (Meunier, 1907) is regarded as the new senior synonym of <em>Sympycnites</em> <em>primaevus</em> Grimaldi &amp; Cumming, 1999, described with a mistaken provenance of Cretaceous Lebanese amber.</p> DANIEL J. BICKEL JOHN MARTIN PRIYA AGNIHOTRI HUKAM SINGH Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 475–486 475–486 10.11646/palaeoentomology.5.5.9 <p><strong>A new species of embidopsocine barklouse in Langhian amber from Zhangpu, China (Psocoptera: Liposcelididae)</strong></p> <p>A new fossil liposcelidid barklouse is described and figured in mid-Miocene amber from southeastern China. <em>Belaphotroctes grimaldii</em> <strong>sp. nov.</strong> is documented from an individual preserved in Langhian amber from Zhangpu, Fujian, China, and is the first fossil species of the family from the Cenozoic of Asia. The species is distinguished from its congeners and comments are provided regarding the implications of a <em>Belaphotroctes</em> Roesler in the Miocene of China.</p> MICHAEL S. ENGEL BO WANG Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 487–492 487–492 10.11646/palaeoentomology.5.5.10 <p><strong><em>Hirsutisoma grimaldii</em> sp. nov., a <em>ca</em>. 99-million-year-old ricinuleid (Primoricinulei, Hirsutisomidae) from Cretaceous Burmese amber with a corticolous, scansorial lifestyle</strong></p> <p>Ricinulei Thorell, 1876 is an order of Arachnida currently represented in the New and Old Worlds by 103 living species. The order is also represented in the fossil record from the Carboniferous (<em>ca</em>. 305–319 Ma) and the Cretaceous (<em>ca</em>. 99 Ma) periods. In the present contribution, <em>Hirsutisoma grimaldii</em> <strong>sp. nov.</strong>, a new extinct species of the suborder Primoricinulei Wunderlich, 2015, is described from a specimen preserved in Cretaceous Burmese amber. The specimen is a well-preserved adult male in which several taxonomically informative structures are visible, allowing the new species to be differentiated from <em>Hirsutisoma bruckschi</em> Wunderlich, 2017, the only other congener for which a male is known. This description raises the number of Cretaceous Ricinulei species to six. A comparative table documents morphological differences among the various species of this lineage. Hypotheses concerning the paleoecology and functional morphology of this species and, by extrapolation, other primoricinuleids, are presented. The evidence suggests that Primoricinulei were corticolous, scansorial predators.</p> RICARDO BOTERO-TRUJILLO STEVEN R. DAVIS PETER MICHALIK LORENZO PRENDINI Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 493–504 493–504 10.11646/palaeoentomology.5.5.11 <p><strong><em>Docidiadia grimaldii</em> sp. nov. from Myanmar amber (Diptera: Diadocidiidae), with comments on the genus <em>Docidiadia</em> Blagoderov &amp; Grimaldi</strong></p> <p><em>Docidiadia grimaldii</em> <strong>sp. nov.</strong>, the second known species in the genus, is described based on a holotype female and on a female paratype. The differences with <em>D. burmitica</em> Blagoderov &amp; Grimaldi, the type-species of the genus, are discussed. The specimens allow the study of important details of the antenna, wing, and terminalia. The question of the connection of <em>Docidiadia</em> to the extant family Diadocidiidae is addressed. An antenna with the flagellum more slender towards the apex is also known in the extinct sciaroid family Archizelmiridae, but the wing venation of <em>Docidiadia</em> is much more conservative than the wing in archizelmirid genera.</p> DALTON de SOUZA AMORIM BRIAN V. BROWN Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 505–512 505–512 10.11646/palaeoentomology.5.5.12 <p><strong>Fossils and flies: Contributions celebrating David A. Grimaldi (Front matter)</strong></p> PHILLIP BARDEN Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 405–405 405–405 <p><strong>The vision of David Grimaldi</strong></p> <p>We are honored to present this special issue of <em>Palaeoentomology</em> in celebration of our dear friend, mentor, and collaborator, David A. Grimaldi (Fig. 1), and on the occasion of his 65<sup>th</sup> birthday on 22 September 2022. This issue has been compiled in recognition of David’s remarkable ongoing impact on the fields of paleontology, entomology, and evolutionary biology. Throughout his career David has worn an inspirational path across disciplines as an exceptional, thoughtful, and creative scientist. His body of work is vast, punctuated by long-lasting classics that are international in scale and recognition. David is a world authority in multiple arenas; his well-earned expertise ranges from the fossil record and evolutionary history of insects beginning 400 million years ago in the Devonian to the systematics and morphology of notoriously complex drosophilids at the species level. His contributions have helped shape modern palaeoentomology. We are grateful for David’s work and fellowship and look forward to his continued accomplishments.</p> PHILLIP BARDEN MICHAEL S. ENGEL Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 406–429 406–429 10.11646/palaeoentomology.5.5.1 <p><strong>The Inventory Imperative</strong></p> <p>“<em>Imagine what could be accomplished—at a fraction of the cost of a Mars rover—by a small army of explorers and taxonomists toiling away to discover and categorize organisms</em>…”—Grimaldi and Engel, 2007</p> <p>“<em>A complete survey of life on Earth may appear to be a daunting task. But compared with what has been dared and achieved in high-energy physics, molecular genetics, and other branches of ‘big science,’ it is in the second or third rank</em>”—E.O. Wilson, 1985</p> QUENTIN WHEELER Copyright (c) 2022 Magnolia press limited 2022-09-22 2022-09-22 5 5 430–433 430–433 10.11646/palaeoentomology.5.5.2