Palaeoentomology <p><strong>Palaeoentomology </strong>is the official journal of the <a href="">International Palaeoentomological Society</a> (IPS). It is an international peer-reviewed scientific journal, which publishes high quality, original research contributions as well as review papers. Papers are published in English and they cover a wide spectrum of topics in palaeoentomology, fossil terrestrial arthropods and amber research, i.e. systematic palaeontology, morphology, diversity, palaeogeography, palaeoecology, palaeobehavior, evolutionary and phylogenetic studies on fossil insects and terrestrial arthropods, biostratigraphy, taphonomy, and amber (deposits, inclusions, geochemistry, curation). Descriptions of new methods (analytical, instrumental or numerical) should be relevant to the broad scope of the journal.</p> <p> </p> <p>Palaeoentomology is the flag journal of IPS, who is responsible for the editing of this journal. For more info about IPS, please contact Prof. Dr. Hab. Dany Azar, Lebanese University, Lebanon.</p> Magnolia press en-US Palaeoentomology 2624-2826 <span lang="EN-GB">Authors need to complete and return an </span><span lang="EN-GB"><a href="/phytotaxa/images/copyright.rtf">Assignment of Copyright</a> </span><span lang="EN-GB">form when a paper is accepted for publication. Authors from institutions that do not allow transfer of copyrights to publishers (e.g. government institutions such as USDA, CSIRO) should attach a copyright waiver or similar document.</span> <strong>Hodotermopsid termites from the mid-Cretaceous Hkamti and Kachin ambers (Isoptera: Hodotermopsidae)</strong> <p>The past diversity of Isoptera stands in stark contrast to their extant diversity as the number of fossil termite species is relatively low. Many early-diverging families are unknown from the Cretaceous, a crucial period encompassing the origins of many extant lineages. Therefore, the study of their past diversity dynamics, leading to their present-day diversity, and origin remains shrouded by a dearth of fossil evidence. Here, we report two new taxa of Hodotermopsidae from the Albian Hkamti and mid-Cretaceous Kachin ambers: <em>Hodotermopsella novella</em> <strong>gen. et sp. nov.</strong> and <em>Tyrannotermes spinifer</em> <strong>gen. et sp. nov.</strong>, the former placed in a new subfamily, Hodotermopsellinae <strong>subfam. nov.</strong> These new species include the earliest occurrences of Hodotermopsidae in the fossil record. We propose new synonymizations for Cretaceous genera from China <em>Jitermes</em> Ren (= <em>Huaxiatermes</em> Ren and <em>Asiatermes</em> Ren, both <strong>syn. nov.</strong>) and <em>Meiatermes</em> Lacasa Ruiz &amp; Martínez-Delclòs (= <em>Caatingatermes</em> Martins-Neto, Ribeiro-Júnior &amp; Prezoto, <em>Araripetermes</em> Martins-Neto, Ribeiro-Júnior &amp; Prezoto, <em>Nordestinatermes</em> Martins-Neto, Ribeiro-Júnior &amp; Prezoto, all <strong>syn. nov.</strong>), and revise the status of the Carinatermitidae <strong>stat. nov.</strong> We discuss the systematic placement of the Pabuonqedidae as well as the implications of the new hodotermopsids on future divergence-time estimates for Teletisoptera and Isoptera.</p> MICHAEL S. ENGEL CORENTIN JOUAULT Copyright (c) 2024 Magnolia press limited 2024-02-27 2024-02-27 7 1 80 91 10.11646/palaeoentomology.7.1.5 <strong>When rare click beetles were not that rare: Cretaceous Cerophytidae Latreille, 1834 (Coleoptera) from Siberia</strong> <p>Cerophytidae, commonly known as rare click beetles, represent one of the more basal lineages of the beetle superfamily Elateroidea, with a rather diverse fossil record in the Mesozoic. Four fossil species of this group were previously described from the Cretaceous of Siberia. Here, we redescribe Aphytocerus communis Zherichin, 1977 based on newly found amber specimens from Yantardakh (Taimyr Peninsula). We were able to locate and herein discuss the type specimens of two Mesozoic Siberian cerophytids, Baissophytum convexum Chang, Kirejtshuk &amp; Ren, 2011 and Baissopsis ampla (Chang, Kirejtshuk &amp; Ren, 2011), which were described from the Lower Cretaceous Zaza Formation in Transbaikalia. We compare them with A. communis and discuss the morphology of all three genera, and with Brachycerophytum cretaceum Yu, Ślipiński &amp; Pang, 2019 from Burmese amber.</p> DMITRY TELNOV EVGENY E. PERKOVSKY DMITRY V. VASILENKO ROBIN KUNDRATA Copyright (c) 2024 Magnolia press limited 2024-02-27 2024-02-27 7 1 92 103 10.11646/palaeoentomology.7.1.6 <strong>Unusual snakefly larvae in about 100 million-year-old amber and the evolution of the larva-pupa transition</strong> <p>The evolutionary success of Insecta, more precisely of its ingroup Holometabola, has partly been explained by their ontogeny, with larvae and adults differing in their morphology and ecology. This differentiation occurs in large and well-known groups such as beetles, butterflies and bees, but also in the relatively species-poor group of snakeflies (Raphidioptera). Despite the rather small number of species, snakeflies are evolutionarily very significant as they were part of the early diversification of Holometabola and still exhibit several plesiomorphic traits retained from the ground pattern of the latter, for example, a mobile pupa. Furthermore, during development, some snakeflies show a mixture of larval and pupal, sometimes even of adult characters, a phenomenon called metathetely. We here report a 100 million-year-old fossil snakefly larva from Myanmar amber with possible characters reminiscent of metathetely. Different dimensions of the body were measured in the specimen and other snakefly larvae and pupae, and ratios were calculated and compared among the larvae. The new fossil shows similarities to extant pupae in the larger length of the prothorax, similarities to modern adults in the small width of the prothorax, but also similarities to other fossil snakefly larvae such as the undivided tarsus and the antenna being subdivided into only five elements. Such a mixture of characters from different developmental stages points to a less pronounced metamorphosis in fossil snakeflies than in extant ones. Similar ontogenetic patterns, with a more gradual development in earlier representatives evolving into a more pronounced metamorphosis in modern representatives, are also known in other groups of Euarthropoda and point to heterochronic events in the evolution of these lineages.</p> JOACHIM T. HAUG ANA ZIPPEL SIMON LINHART PATRICK MÜLLER CAROLIN HAUG Copyright (c) 2024 Magnolia press limited 2024-02-27 2024-02-27 7 1 104 111 10.11646/palaeoentomology.7.1.7 <strong>First record of insects from the oldest and older Dryas of Altai (Russia). Coleoptera assemblages from Lebed River</strong> <p>Subfossil remains of insects found in the Lebed site (52.25220°N, 87.15692°E) located on the Lebed River, Altai Republic are recorded. The calibrated radiocarbon dates for two layers of these deposits were 16,461–17,056 cal BP, and 13,520–14,077 cal BP, which correspond to the oldest and older Dryas (Late Pleistocene). Insect assemblages of these deposits are mainly represented by Coleoptera, which are noteworthy there for high taxonomic and ecological diversity and include several endemic and relic species. At least 120 beetle species from 17 families have been found altogether, of them, 37 species are recorded for the Pleistocene deposits of Western Siberia for the first time. Three families, Carabidae, Staphylinidae and Curculionidae are most abundantly represented in the examined Lebed site. Ecologically, this beetle assemblage is dominated by species that are currently confined to the taiga belt and alpine meadows of the Altai Mountains; aquatic and near-water species are also well represented. The studied assemblages are rather different from the previously studied Pleistocene insect fauna in the south of the West Siberian Plain. The taxonomic and ecological compositions of the beetle fauna of Lebed site suggest its existence under humid palaeoclimate that was significantly colder than modern climate in this area.</p> ANNA A. GURINA ROMAN Yu. DUDKO YURI E. MIKHAILOV ALEXANDER A. PROKIN ALEXEY Yu. SOLODOVNIKOV EVGENII V. ZINOVYEV ANDREI A. LEGALOV Copyright (c) 2024 Magnolia press limited 2024-02-27 2024-02-27 7 1 112 131 10.11646/palaeoentomology.7.1.8 <strong>A new Triassic Tettigarctidae (Insecta, Hemiptera) from the Amisan Formation (Republic of Korea)</strong> <p>A new species of Tettigarctidae Sanmai? zetavena sp. nov. is described and illustrated from the Late Triassic Amisan Formation (Republic of Korea). This new species is the oldest representative of the genus Sanmai Chen et al., 2016, previously only known from the Middle Jurassic of China. This discovery expands the temporal range of the genus Sanmai and increases the morphological diversity of Cicadomorpha during the Triassic. Together with recent discoveries of new fossil insect taxa, the description of this new species advocates for further study of the Amisan Formation palaeoentomofauna.</p> CORENTIN JOUAULT GI-SOO NAM MATHIEU BODERAU SEUNG-HYUK KWON ANDRÉ NEL Copyright (c) 2024 Magnolia press limited 2024-02-27 2024-02-27 7 1 132 140 10.11646/palaeoentomology.7.1.9 <strong>A new species of long-legged flies with a brief review and key to species of the genus <em>Wheelerenomyia</em> Meunier from Eocene Baltic amber (Diptera: Dolichopodidae)</strong> <p>A new species <em>Wheelerenomyia negrobovi</em> <strong>sp. nov.</strong>, is described and illustrated from the late Eocene Baltic amber. The genus is placed in the extant subfamily Sciapodinae and includes 13 recognized species, all from Baltic amber. A key to species of this genus is proposed. The following new synonyms are proposed:<em> Wheelerenomyia</em> <em>eocenica</em> Meunier, 1907 = <em>Nematoproctus subparvus </em>Meunier, 1916, <strong>syn. nov</strong>.; <em>Wheelerenomyia</em> <em>originaria</em> (Meunier, 1907) = <em>Amesorhaga quadrispinosa </em>Negrobov &amp; Selivanova, 2003, <strong>syn. nov</strong>.; <em>Wheelerenomyia parva</em> (Meunier, 1907) = <em>Nematoproctus parvulus </em>Meunier, 1907, <strong>syn. nov</strong>.</p> IGOR YA. GRICHANOV Copyright (c) 2024 Magnolia press limited 2024-02-27 2024-02-27 7 1 141 147 10.11646/palaeoentomology.7.1.10 <strong>Supplement to the Burmese (Myanmar) amber checklist and bibliography, 2023</strong> <p>This is a supplement to the Burmese (Myanmar) amber checklist and bibliography covering taxa described or recorded during 2023, plus a couple of earlier records that were missed previously. Up to the end of 2023, 2,781 species have been recorded from Kachin amber, of which 244 were named in 2023; 16 species have been recorded from older Hkamti amber (six were named in 2023), of which two are known from both Hkamti and Kachin amber. Another four species were named in 2023, though it is uncertain whether they are in Kachin or Hkamti amber. In total 253 species were named from Cretaceous amber from Myanmar in 2023.</p> ANDREW J. ROSS Copyright (c) 2024 Magnolia press limited 2024-02-27 2024-02-27 7 1 148 165 10.11646/palaeoentomology.7.1.11 <strong>A new fossil genus near <em>Meghyperus</em> Loew from Baltic amber (Diptera: Atelestidae)</strong> <p>The Atelestidae are a small family of empidoid flies (Diptera: Empidoidea), including 15 recent species in five genera (Sinclair &amp; Grimaldi, 2020). In contrast, they have a rather high fossil diversity (Table 1), with 28 species in nine genera. These fossil taxa are almost entirely from Cretaceous ambers (Canadian, Lebanese, Myanmar, New Jersey, Spanish), and prior to this study only a single described species from Baltic amber (Nemedina eocenica Sinclair &amp; Arnaud, 2001) was known. The extant species are widespread, with two Afrotropical, three Neotropical, two Nearctic and eight Palaearctic species. This family is recognized within the Empidoidea on the basis of their well-developed anal lobe of the wing, distinct alula, R4+5 unforked, M1+2 usually unforked or forked beyond cell dm, cell cua long, at least as long as cell bm, female tergite 10 absent, male terminalia symmetrical and unrotated, with elongate gonocoxal apodemes and shortened hypandrium (Chvála, 1983; Grimaldi &amp; Cumming, 1999; Sinclair &amp; Cumming, 2006).</p> BRADLEY J. SINCLAIR ANDREAS STARK CHRISTEL HOFFEINS AGNIESZKA SOSZYŃSKA Copyright (c) 2024 Magnolia press limited 2024-02-27 2024-02-27 7 1 68 71 10.11646/palaeoentomology.7.1.2 <strong>A new species of dustywings (Neuroptera: Coniopterygidae) from mid-Cretaceous Kachin amber</strong> <p>speciesThe family Coniopterygidae, commonly known as dustywings, occupies a distinctive niche within the Neuroptera, as they are the smallest members of this order (Grimaldi &amp; Engel, 2005; Engel <em>et al</em>., 2018). Characterized by forewing lengths seldom surpassing 5 mm, with a few exceptions (Liu &amp; Lu, 2017), this family has achieved a remarkable speciosity, with approximately 570 extant species scattered across diverse habitats globally (Oswald &amp; Machado, 2018).</p> CORENTIN JOUAULT MICHAEL S. ENGEL Copyright (c) 2024 Magnolia press limited 2024-02-27 2024-02-27 7 1 72 75 10.11646/palaeoentomology.7.1.3 <strong>SEM morphological study on carapace of <em>Cyclestheria hislopi </em>and comparison with fossil taxa</strong> <p>The circum-tropically distributed cryptic extant clam shrimp species Cyclestheria hislopi (Baird, 1859) is thought to be the only species of Cyclestherida, although molecular data show that its populations on different continents have large genetic distance (Schwentner et al., 2013). Its parthenogenetical reproduction and direct ovoviviparous development in the dorsal brood chamber indicate that C. hislopi is the sister taxon of all Cladocera (Olsen et al., 1996; Olsen, 1999; Olesen &amp; Richter, 2013; Schwentner et al., 2018). While the large carapace, capable of enclosing the whole body, shows a close similarity with spinicaudatans. In this paper we document its carapace ornamentation by the use of a scanning electron microscope (SEM) and compare to related fossil taxa.</p> GANG LI SHUSEN SHU Copyright (c) 2024 Magnolia press limited 2024-02-27 2024-02-27 7 1 76 79 10.11646/palaeoentomology.7.1.4 <strong>Time-traveling through fossil planthopper tegmina in the Paleozoic and Mesozoic eras (Insecta: Hemiptera: Fulgoromorpha)</strong> <p>This study presents a comprehensive and illustrated catalogue of the forewing venation of the Paleo- and Mesozoic Hemiptera planthoppers. It compiles fragmented information dispersed across various publications, ensuring a uniform interpretation of tegmina venation for fossil taxa. Taxa are presented by family and each species is documented with its stratigraphic age, and tegmina are redrawn from their original illustrations and reinterpreted with relevant comments when necessary. This catalogue aims to be a decisive tool for research into the taxonomy and phylogeny of the group by providing foundational data for comparative analyses of forewing patterns in Fulgoromorpha, as it represents a valuable shared dataset for both fossil and extant taxa. Yanducixius is formally placed in Lalacidae.</p> MANON BUCHER GODEFROI GIGNOUX JACEK SZWEDO THIERRY BOURGOIN Copyright (c) 2024 Magnolia press limited 2024-02-27 2024-02-27 7 1 1 67 10.11646/palaeoentomology.7.1.1