Palaeoentomology <p><strong>Palaeoentomology </strong>is the official journal of the <a href="">International Palaeoentomological Society</a> (IPS). It is an international peer-reviewed scientific journal, which publishes high quality, original research contributions as well as review papers. Papers are published in English and they cover a wide spectrum of topics in palaeoentomology, fossil terrestrial arthropods and amber research, i.e. systematic palaeontology, morphology, diversity, palaeogeography, palaeoecology, palaeobehavior, evolutionary and phylogenetic studies on fossil insects and terrestrial arthropods, biostratigraphy, taphonomy, and amber (deposits, inclusions, geochemistry, curation). Descriptions of new methods (analytical, instrumental or numerical) should be relevant to the broad scope of the journal.</p> <p> </p> <p>Palaeoentomology is the flag journal of IPS, who is responsible for the editing of this journal. For more info about IPS, please contact Prof. Dr. Hab. Dany Azar, Lebanese University, Lebanon.</p> en-US <span lang="EN-GB">Authors need to complete and return an </span><span lang="EN-GB"><a href="/phytotaxa/images/copyright.rtf">Assignment of Copyright</a> </span><span lang="EN-GB">form when a paper is accepted for publication. Authors from institutions that do not allow transfer of copyrights to publishers (e.g. government institutions such as USDA, CSIRO) should attach a copyright waiver or similar document.</span> (Diying Huang) (Journal support team) Thu, 23 Jun 2022 00:00:00 +1200 OJS 60 <p><strong>A peculiar new genus of Scytinopteridae (Hemiptera, Cicadomorpha) from the Permian-Triassic boundary beds of Mongolia</strong></p> <p>Scytinopteridae Handlirsch, 1906 is a widespread family of extinct Cicadomorpha known worldwide from the Permian and Triassic (Lambkin, 2016). This family was understood very broadly (<em>e.g.</em>, Evans, 1956), but was later restricted to the genera with heavily sclerotized punctate tegmina having a costal fracture and a hypocostal socket fixed on the thorax in repose, and was placed in the superfamily Scytinopteroidea Handlirsch, 1906 (Shcherbakov, 1984). Based on these features, scytinopteroids are considered inhabitants of waterside vegetation capable of temporary submergence, and are ancestors of true bugs (Shcherbakov, 1996, 2000). In the beds near the Permian-Triassic boundary of Mongolia, an unusual new genus of Scytinopteridae was discovered with pointed coleopterous tegmina lacking the costal fracture and the claval furrow. These modifications indicate that the flying ability in the new scytinopterid was reduced or lost, and the peculiar shape of the tegmina probably helped to mimic host plant buds or seeds (plant-part mimicry).</p> DMITRY E. SHCHERBAKOV Copyright (c) 2022 Magnolia press limited Thu, 23 Jun 2022 00:00:00 +1200 <p><strong>Debris-carrying psocodean nymph from Lebanese amber</strong></p> <p>Camouflage, or decorating, has been reported in fossilised immatures of different insect groups, notably Neuroptera (Chrysopidae, Nymphidae, and Ascalaphidae), Coleoptera (Chrysomelidae), Trichoptera, Lepidoptera, Hemiptera (Gelastocoridae, Reduviidae), and Psocodea (‘Psocoptera’ or non-parasitic lice) (Pérez-de la Fuente <em>et al.</em>, 2012; Wang <em>et al.</em>, 2016; Kiesmüller <em>et al.</em>, 2021; Xu <em>et al.</em>, 2021). Such behaviours allow effective concealment of immatures from predators and/or prey by carrying vegetal debris, soil, dust and, sometimes, arthropod remains on the thorax and/or abdomen.</p> MARINA HAKIM, DI-YING HUANG, DANY AZAR Copyright (c) 2022 Magnolia press limited Thu, 23 Jun 2022 00:00:00 +1200 <p><strong>Edmund Jarzembowski at 70: An appreciation</strong></p> <p>Edmund Aleksander Jarzembowski (BSc PhD FGS FRES) is currently a Leverhulme Emeritus Fellow; Scientific Associate (researcher) at The Natural History Museum London (NHMUK); and Professor at the Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences (NIGPAS), Nanjing, specializing in the study of fossil insects (palaeoentomology).</p> PETER A. AUSTEN, BO WANG, ANDREW J. ROSS, ROBERT A. CORAM Copyright (c) 2022 Magnolia press limited Thu, 23 Jun 2022 00:00:00 +1200 <p><strong>A new caddisfly of the family Baissoferidae (Insecta: Trichoptera) from the Lower Cretaceous of Mongolia</strong></p> <p>This article is devoted to the description of a new representative of the family Baissoferidae from the Khutel-Khara locality (Mongolia, East Gobi Aimag) located 75 km southwest of Mount Sain-Shand. The insect-bearing deposits of the Tsagan Tsab Formation, the Lower Tsagan Tsab Subformation in this locality are dated to the earliest Cretaceous, Berriasian-Valanginian. In total, about 3,300 fossil insect imprints of 14 orders were collected from the Khutel-Khara locality. It is important that the fauna of insects has both Jurassic and Cretaceous features. For example, many mayfly and stonefly nymphs that lived there (Sinitshenkova, 1987, 1989) were still typical Jurassic lacustrine forms, while among terrestrial insects, on the contrary, forms that occur mainly in the Lower Cretaceous of Transbaikalia (some rove beetles and scarabaeoids) prevailed (Ponomarenko, 2021). It is especially interesting that many forms are represented in Khutel-Khara, first appearing in the geological record, then widespread in the Lower Cretaceous. This primarily concerns hymenopteran insects (Rasnitsyn, 1990; Kopylov &amp; Rasnitsyn, 2020).</p> IRINA D. SUKATSHEVA, NINA D. SINITSHENKOVA Copyright (c) 2022 Magnolia press limited Thu, 30 Jun 2022 00:00:00 +1200 <p>The oldest fossil of the sawfly subfamily Nematinae (Hymenoptera: Tenthredinidae) from the Paleocene of Menat (France)</p> <p>The speciose family Tenthredinidae has a scarce fossil record (Nel <em>et al.</em>, 2022a, b), despite recent molecular-based phylogenetic analyses suggesting a Cretaceous age for the origin (<em>ca</em>. 100 Ma) and the diversification of the family into its modern subfamilies (Nyman <em>et al.</em>, 2006, 2019; Niu <em>et al.</em>, 2021). The lack of Mesozoic representative of this group remains unexplained because Nyman <em>et al</em>. (2006) dated its origin at 100 Ma. Cenozoic fossils are currently attributed to the subfamilies Allantinae, Heterarthrinae, Blennocampinae, and Nematinae (Taeger <em>et al</em>., 2010; Nel <em>et al</em>, 2022a, b).</p> ANDRÉ NEL Copyright (c) 2022 Magnolia press limited Thu, 30 Jun 2022 00:00:00 +1200 <p><strong><em>Cretoneuronema</em> gen. nov. (Neuroptera: Hemerobiidae), a new brown lacewing genus from the mid-Cretaceous Kachin amber</strong></p> <p>A new genus and species of the lacewing family Hemerobiidae, <em>Cretoneuronema jarzembowskii</em> <strong>gen. et sp. nov.</strong>, is described from the mid-Cretaceous Kachin amber in northern Myanmar. This new genus has a definite hemerobiid affinity and is characterized by the distally fused ScP and RA in both fore- and hind wing, the presence of forewing RP and MA (= ORB1 and ORB2) separately diverging from R near wing base and closely spaced with each other, the forewing MA with anteriorly directed branches, the deeply forked forewing CuP, and the completely developed hind wing CuP. The new genus resembles some extinct brown lacewing genera, such as <em>Promegalomus </em>Panfilov, 1980, <em>Cretomerobius</em> Ponomarenko, 1992, and <em>Proneuronema </em>Makarkin, Wedmann &amp; Weiterschan, 2016, from the Late Jurassic to Eocene, and represents the second Kachin amber genus of Hemerobiidae, a family that appears to be very rare from this deposit.</p> XING-YUE LIU, ZU-LUAN CHEN, DE ZHUO Copyright (c) 2022 Magnolia press limited Thu, 23 Jun 2022 00:00:00 +1200 <p><strong>An enigmatic Tettigoniidea from the Lower Cretaceous amber of Bqaatouta, Lebanon (Orthoptera, Ensifera)</strong></p> <p><em>Aenigmaraphidophora</em> <em>mouniri</em> <strong>gen. et sp. nov.</strong>, an enigmatic representative of the ensiferan infraorder Tettigoniidea, is described and figured from the Lower Cretaceous of Lebanon, based on a nearly complete male specimen. The new fossil could fit in either in the stem group of the Rhaphidophoridae or in the Tettigoniidae. In either cases, the discovery is congruent with the current hypotheses on the dating of these families. We discuss on the consequence of the presence of an external fore tibial tympan in the new fossil, while absent in the extant Rhaphidophoridae.</p> DANY AZAR, RAMY MAALOUF, ANDRÉ NEL Copyright (c) 2022 Magnolia press limited Thu, 23 Jun 2022 00:00:00 +1200 <p><strong>The second species of <em>Rudiaeschna</em> (Odonata, Rudiaeschnidae) discovered in the Lower Cretaceous of Inner Mongolia, Northeast China</strong></p> <p><em>Rudiaeschna jarzembowskii</em> <strong>sp. nov.</strong>, the second species of the small aeshnopteran family Rudiaeschnidae, is described from the Lower Cretaceous Yixian Formation at the Liutiaogou locality, Ningcheng County, Inner Mongolia, NE China. The new species differs from the type species of the family, namely <em>Rudiaeschna limnobia,</em> in possessing less cells and crossveins in nearly all parts of forewing. It also shows a distally forked vein RP2, a character that was previously only known in taxa of the much more recent and derived aeshnopteran family Aeshnidae.</p> DI-YING HUANG, JOUAULT CORENTIN, ANDRÉ NEL Copyright (c) 2022 Magnolia press limited Mon, 27 Jun 2022 00:00:00 +1200 <p><strong><em>Homalenchodes</em>, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae)</strong></p> <p>A new fossil genus and species of the family Melandryidae, <em>Homalenchodes</em> <em>jarzembowskii</em> Li, Hsiao, Yoshitomi &amp; Cai <strong>gen. et sp. nov.</strong>, is described and illustrated from the mid-Cretaceous Burmese amber. <em>Homalenchodes</em> is likely closely related to the serropalpin genus <em>Enchode</em>s LeConte, based on their simple maxillary palps, complete prosternal process, exposed protrochantins, ventrally lobed penultimate metatarsomeres, and absence of transverse rows of bristles or spines along the meso- and metatibiae.</p> YAN-DA LI, YUN HSIAO, HIROYUKI YOSHITOMI, DI-YING HUANG, CHEN-YANG CAI Copyright (c) 2022 Magnolia press limited Mon, 27 Jun 2022 00:00:00 +1200 <p><strong>Revision of <em>Archizelmira</em> Rohdendorf, 1962 (Diptera, Archizelmiridae)</strong></p> <p>A new species of the extinct genus <em>Archizelmira</em> Rohdendorf, 1962 is described based on an isolated wing from the Middle Jurassic of West Siberia (Itat Formation, Bathonian). Both previously known species are re-examined. Additional wing venation characters are found for the genus, which support its status as a sister group to the rest of the family. A new Cretaceous subfamily, Burmazelmirinae <strong>subfam. nov.</strong> is proposed.</p> ELENA DMITRIEVNA LUKASHEVICH Copyright (c) 2022 Magnolia press limited Wed, 29 Jun 2022 00:00:00 +1200 <p><strong><em>Paleopsychoda jarzembowskii</em>, a new Lower Cretaceous species of moth flies from Lebanese amber (Diptera: Psychodidae: Psychodinae)</strong></p> <p><em>Paleopsychoda jarzembowskii </em><strong>sp. nov.</strong> is characterized, illustrated, and described from the lower Barremian amber of Mdeyrij-Hammana (Lebanon). It represents the first psychodid fly in Lower Cretaceous amber showing sexual dimorphism in wing characters. The discovery of this Cretaceous psychodid fly demonstrates that this genus had a large diversification and broad distribution, and improves our knowledge of the evolution of moth flies.</p> DANY AZAR, SIBELLE MAKSOUD Copyright (c) 2022 Magnolia press limited Wed, 29 Jun 2022 00:00:00 +1200 <p>The oldest crown representative of the family Aulacidae (Hymenoptera: Evanioidea) from the Paleocene of Menat (France)</p> <p><em>Pristaulacus</em> <em>jarzembowskii</em> <strong>sp. nov.</strong>, oldest fossil of the crown group Aulacidae, is described from the Paleocene of Menat (France). The previously oldest fossil crown-aulacid was known from the Lowermost Eocene amber of Oise (France). The inferences of parasitoidism on Cerambycidae or Buprestidae together with the preferences of warm evergreen forest with wildfires are in accordance with the known data on the Menat paleobiota.</p> CORENTIN JOUAULT, ANDRÉ NEL Copyright (c) 2022 Magnolia press limited Wed, 29 Jun 2022 00:00:00 +1200 <p><strong>Another strange holometabolan larva from Kachin amber—the enigma of the beak larva (Neuropteriformia)</strong></p> <p>Holometabolan larvae are dominating components of modern terrestrial and freshwater ecosystems and have a significant ecological impact. Also in past ecosystems, various types of such larvae have been present, which is especially well known from ambers from all over the world. During the Cretaceous, holometabolan larvae with a very modern appearance co-occur with those of morphologies totally unknown in the ecosystems of today. One of these morphologies only known from <em>ca</em>. 100-million-year-old Kachin amber from Myanmar is represented by the so-called “beak larvae”, which possess an anteriorly projecting beak-like mouth cone, previously being described from two specimens. We describe here a third specimen as a new species, ?<em>Partisaniferus edjarzembowskii</em> <strong>sp. nov.</strong> This new species differs from the previously described beak larva species <em>Partisaniferus atrickmuelleri</em> in the shape of the trunk end as well as in lacking a differentiation of the tergites into distinct sclerites and in the absence of abdomen protrusions. We discuss possible aspects of the ontogeny of the beak larvae, including the possibility that the here described specimen and one of the previously known ones are different larval stages of ?<em>P. edjarzembowskii</em> <strong>sp. nov.</strong> Furthermore, we discuss possible relationships of beak larvae within Neuropteriformia.</p> JOACHIM T. HAUG, CAROLIN HAUG Copyright (c) 2022 Magnolia press limited Thu, 30 Jun 2022 00:00:00 +1200