Abstract
A new Epidendrum species of the Nocturnum group is proposed for the Brazilian central Amazon, near Manaus. It is described, illustrated, and compared with similar species. Epidendrum dayseae can be recognized by the pendent stem, the long, narrow leaves, a relatively long floral pedicel, and the lateral lobes of the lip smaller than the mid-lobe and deeply separated. The new species resembles E. longicolle, but is distinguished by the union between of the lateral and mid-lobes of the lip being under half the mid-lobe length. The new species is also compared with E. plurifolionocturnum. Its chromosome number is 2n = 4x = 80, with a band pattern similar to other species of the Nocturnum group.
References
<p>Assis, F.N.M., Souza, B.C.Q., Medeiros-Neto, E., Pinheiro, F., Barros, E., Silva, A.E. & Felix, L.P. (2013) Karyology of the genus <em>Epidendrum</em> (Orchidaceae: Laeliinae) with emphasis on subgenus <em>Amphiglottium</em> and chromosome number variability in <em>Epidendrum secundum</em>. <em>Botanical Journal of the Linnaean Society</em> 172 (3): 329–44. https://doi.org/10.1111/boj.12045</p>
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<p>Carnevali, G., Ramírez-Morillo, I.M., Romero-González, G.A., Vargas, C.A. & Foldats, E. (2003) Orchidaceae. <em>In</em>: Berry, P.E., Yatskievych, K. & Holts, B.K. (Eds.) <em>Flora of the Venezuelan Guyana</em> 7: 200–618.</p>
<p>Cordeiro, J.M., Kaehler, M., Souza, L.G. & Felix, L.P. (2020) Heterochromatin and numeric chromosome evolution in Bignoniaceae, with emphasis on the Neotropical clade <em>Tabebuia</em> alliance. <em>Genetics and Molecular Biology</em> 43 (1): e20180171. https://doi.org/10.1590/1678-4685-gmb-2018-0171</p>
<p>Cordeiro, J.M.P. (2019) <em>Citotaxonomia do gênero neotropical Epidendrum L. (Laeliinae, Orchidaceae): discutindo grupos infragenéricos.</em> (Thesis). Doctorate in Agronomy, Federal University of Paraiba, 122 pp.</p>
<p>Dressler, R.L. (1993) <em>Phylogeny and Classification of the Orchid Family. </em>Dioscorides Press, Portland, 314 pp.</p>
<p>Dunsterville, G.C.K. & Garay, L.A. (1959) <em>Venezuelan Orchids Illustrated</em> <em>[Dunsterville & Garay] 1</em>. Andre Deutsch, London. 447 pp.</p>
<p>Dunsterville, G.C.K. & Garay, L.A. (1961) <em>Venezuelan Orchids Illustrated</em> <em>[Dunsterville & Garay] 2</em>. Andre Deutsch, London. 360 pp.</p>
<p>Dunsterville, G.C.K. & Garay, L.A. (1965) <em>Venezuelan Orchids Illustrated</em> <em>[Dunsterville & Garay] 3</em>. Andre Deutsch, London. 348 pp</p>
<p>Dunsterville, G.C.K. & Garay, L.A. (1966) <em>Venezuelan Orchids Illustrated</em> <em>[Dunsterville & Garay] 4</em>. Andre Deutsch, London. 463 pp.</p>
<p>Dunsterville, G.C.K. & Garay, L.A. (1972) <em>Venezuelan Orchids Illustrated</em> <em>[Dunsterville & Garay] 5</em>. Andre Deutsch, London. 334 pp.</p>
<p>Dunsterville, G.C.K. & Garay, L.A. (1976) <em>Venezuelan Orchids Illustrated</em> <em>[Dunsterville & Garay] 6</em>. Andre Deutsch, London. 463 pp.</p>
<p>Felix, L.P. & Guerra, M. (2010) Variation in chromosome number and the basic number of subfamily Epidendroideae (Orchidaceae). <em>Botanical Journal of the Linnaean Society</em> 163 (2): 234–278. https://doi.org/10.1111/j.1095-8339.2010.01059.x</p>
<p>Guerra, M. (1986) Reviewing the chromosome nomenclature of Levan <em>et al.</em> <em>Revista Brasileira de Genética</em> 9: 741–743.</p>
<p>Hágsater, E. (1984) Towards an Understanding of the Genus <em>Epidendrum</em>. <em>In:</em> Tan, K.W. (Ed.) <em>Proceedings of the Eleventh World Orchid Conference, Miami, U.S.A.</em> pp. 195–201.</p>
<p>Hágsater, E. & Krahl, A.H. (2020) <em>Epidendrum deditae</em>, a new species of Orchidaceae (Laeliinae) from northern Brazil. <em>Phytotaxa </em>440 (3): 225–231. https://doi.org/10.11646/phytotaxa.440.3.4</p>
<p>Hágsater, E. & Salazar, G.A. (1993) The genus<em> Epidendrum, </em>Part 1. A century of new species in<em> Epidendrum</em>. <em>Icones Orchidacearum 2</em>: pl. 101–200.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2018) <em>Epidendrum plurifolionocturnum</em>. <em>In:</em> Hágsater, E. & Santiago, E. (Eds.) The genus <em>Epidendrum</em>, Part 13. <em>Icones Orchidacearum</em> 16 (2): pl. 1695.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2001) The genus <em>Epidendrum</em>, Part 3. A third century of new species in <em>Epidendrum</em>.<em> Icones Orchidacearum </em>4: pl. 401–500.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2004) The genus <em>Epidendrum</em>, Part 4. A fourth century of new species in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>7: pl. 701–800.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2006) The Genus<em> Epidendrum</em>, Part 5. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>8: pl. 801–900.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2007) The genus <em>Epidendrum</em>, Part 6. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>9: pl. 901–1000.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2008) The genus <em>Epidendrum</em>, Part 7. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>11: pl. 1101–1200.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2009) The genus <em>Epidendrum</em>, Part 8. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>12: pl. 1201–1300.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2010) The genus <em>Epidendrum</em>, Part 9. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>13: pl. 1301–1400.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2013) The genus <em>Epidendrum</em>, Part 10. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>14: pl. 1401–1500.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2015) The Genus <em>Epidendrum</em>, Part 11. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>15 (1): pl. 1501–1568.</p>
<p>Hágsater, E. & Santiago, E. (2018a) The genus <em>Epidendrum</em>, Part 12. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>16 (1): pl. 1601–1667.</p>
<p>Hágsater, E. & Santiago, E. (2018b) The genus <em>Epidendrum</em>, Part 12. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum</em> 16 (2): pl. 1668–1700.</p>
<p>Hágsater, E. & Santiago, E. (2019) The genus <em>Epidendrum</em>, Part 12. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>17 (1): pl. 1701–1756.</p>
<p>Hágsater, E. & Santiago, E. (2020a) The genus <em>Epidendrum</em>, Part 13. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>17 (2): pl. 1757–1800.</p>
<p>Hágsater, E. & Santiago, E. (2020b) The genus <em>Epidendrum</em>, Part 14. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>18 (1): pl. 1849–1848.</p>
<p>Hágsater, E. & Santiago, E. (2021) The genus <em>Epidendrum</em>, Part 14. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>18 (2): pl. 1801–1900.</p>
<p>Hágsater, E. & Soto-Arenas, M. (2005a) <em>Epidendrum</em>. <em>In</em>: Pridgeon, A.M., Cribb, P.J., Chase, M.W. & Rasmussen, F.N. (Eds.) <em>Genera Orchidacearum 4: Epidendroideae (Part 1)</em>. Oxford University Press, Oxford, pp. 236–251.</p>
<p>Hágsater, E. & Soto-Arenas, M. (2005b) Towards an understanding of diversification of the genus <em>Epidendrum. In: </em>Raynal-Roques, A., Roquenant, A. & Prat, D. (Eds.) <em>Proceedings of the 18th World Orchid Conference</em>, Dijon France, Naturalia, Abstract. pp. 225.</p>
<p>Hágsater, E., Archila, F. & Chiron, G. (2015) <em>Epidendrum ixilum</em>. <em>In:</em> Hágsater, E. & Santiago, E. (Eds.) The Genus <em>Epidendrum</em>, Part. 11. <em>Icones Orchidacearum </em>15 (1): pl. 1529.</p>
<p>Hágsater, E., Sánchez, L.S. & García-Cruz, J. (1999) The genus <em>Epidendrum</em>, Part 2. A second century of new species in <em>Epidendrum</em>. <em>Icones Orchidacearum</em> 3: pl. 301–400.</p>
<p>Lindley, J. (1834) <em>Epidendrum longicolle</em>. <em>Edwards’s Botanical Register </em>24 (Misc. 24): 49.</p>
<p>Linnaeus, C. (1759) <em>Epidendrum ciliare</em>. <em>Systema Naturae, Editio Decima </em>2: 1246.</p>
<p>Linnaeus, C. (1763) <em>Species Plantarum, ed. 2, vol. 2</em>. Imprensis Direct. Laurentii Salvii, Holmiae, 899 pp.</p>
<p>Moosburg, M., Hágsater, E. & Ochse, M. (2014) Ökologische Zusammenhänge anhand von Sammlungsexemplaren erkennen: Adhemarius sexoculata Grote, 1867, als Bestäuber bei der Orchideengattung <em>Epidendrum</em> Linnaeus, 1753 (Lepidoptera: Sphingidae). <em>Entomologische Zeitschrift, Stuttgart</em> 124: 2013–2014.</p>
<p>Mori, S.A., Silva, L.A., Lisboa, G. & Coradin, L. (1989) <em>Manual de Manejo do Herbário Fanerogâmico</em>. Ceplac, Ilhéus, 104 pp.</p>
<p>Pessoa, E.M. (2021) <em>Epidendrum</em>. <em>In:</em> Flora do Brasil 2020, Jardim Botânico do Rio de Janeiro, Brasil. Available from: http://floradobrasil.jbrj.gov.br/ (accessed 20 April 2021)</p>
<p>Pessoa, E.M., Cordeiro, J.M.P., Felix, L.P., Almeida, E.M., Costa, L., Nepomuceno, Á., Souza, G., Chase, M., Alves, M. & Van den Berg, C. (2020) Too many species: morphometrics, molecular phylogenetics and genome structure of a Brazilian species complex in <em>Epidendrum</em> (Laeliinae; Orchidaceae) reveal fewer species than previously thought. <em>Botanical Journal of the Linnaean Society</em> 195 (2): 161–188. https://doi.org/10.1093/botlinnean/boaa071</p>
<p>Salazar, G.A. & Dressler, R.L. (2011) The leaves got it right again: DNA phylogenetics supports a sister-group relationship between <em>Eurystyles </em>and<em> Lankesterella </em>(Orchidaceae: Spiranthinae). <em>Lankesteriana </em>11 (3): 337–347. https://doi.org/10.15517/lank.v11i3.18289</p>
<p>Sambin, A. & Aucourd, M. (2021) Mise à jour de l’inventaire des orchidées présentes autour et sur le pic Coudreau des Monts Bakra. Avec une note taxinomique pour quelques espèces. <em>Richardiana N.S. </em>5: 121–141.</p>
<p>Sambin, A. & Essers, D. (2018) Un nouvel <em>Epidendrum</em> (Orchidaceae) du groupe «Nocturnum» de Guyane. <em>Richardiana N.S. </em>2: 144–151.</p>
<p>Schneider, C.A., Rasband, W.S. & Eliceiri, K.W. (2012) NIH Image to ImageJ: 25 years of image analysis. <em>Nature Methods </em>9: 671–675. https://doi.org/10.1038/nmeth.2089</p>
<p>Schweinfurth, C. (1959) Orchids of Peru. <em>Fieldiana Botany</em> 30: 261–531.</p>
<p>Cameron, K.M. (2005) Leave it to the leaves: a molecular phylogenetic study of Malaxideae (Epidendroideae, Orchidaceae).<em> American Journal of Botany </em>92 (6): 1025–1032. https://doi.org/10.3732/ajb.92.6.1025</p>
<p>Carnevali, G., Ramírez-Morillo, I.M., Romero-González, G.A., Vargas, C.A. & Foldats, E. (2003) Orchidaceae. <em>In</em>: Berry, P.E., Yatskievych, K. & Holts, B.K. (Eds.) <em>Flora of the Venezuelan Guyana</em> 7: 200–618.</p>
<p>Cordeiro, J.M., Kaehler, M., Souza, L.G. & Felix, L.P. (2020) Heterochromatin and numeric chromosome evolution in Bignoniaceae, with emphasis on the Neotropical clade <em>Tabebuia</em> alliance. <em>Genetics and Molecular Biology</em> 43 (1): e20180171. https://doi.org/10.1590/1678-4685-gmb-2018-0171</p>
<p>Cordeiro, J.M.P. (2019) <em>Citotaxonomia do gênero neotropical Epidendrum L. (Laeliinae, Orchidaceae): discutindo grupos infragenéricos.</em> (Thesis). Doctorate in Agronomy, Federal University of Paraiba, 122 pp.</p>
<p>Dressler, R.L. (1993) <em>Phylogeny and Classification of the Orchid Family. </em>Dioscorides Press, Portland, 314 pp.</p>
<p>Dunsterville, G.C.K. & Garay, L.A. (1959) <em>Venezuelan Orchids Illustrated</em> <em>[Dunsterville & Garay] 1</em>. Andre Deutsch, London. 447 pp.</p>
<p>Dunsterville, G.C.K. & Garay, L.A. (1961) <em>Venezuelan Orchids Illustrated</em> <em>[Dunsterville & Garay] 2</em>. Andre Deutsch, London. 360 pp.</p>
<p>Dunsterville, G.C.K. & Garay, L.A. (1965) <em>Venezuelan Orchids Illustrated</em> <em>[Dunsterville & Garay] 3</em>. Andre Deutsch, London. 348 pp</p>
<p>Dunsterville, G.C.K. & Garay, L.A. (1966) <em>Venezuelan Orchids Illustrated</em> <em>[Dunsterville & Garay] 4</em>. Andre Deutsch, London. 463 pp.</p>
<p>Dunsterville, G.C.K. & Garay, L.A. (1972) <em>Venezuelan Orchids Illustrated</em> <em>[Dunsterville & Garay] 5</em>. Andre Deutsch, London. 334 pp.</p>
<p>Dunsterville, G.C.K. & Garay, L.A. (1976) <em>Venezuelan Orchids Illustrated</em> <em>[Dunsterville & Garay] 6</em>. Andre Deutsch, London. 463 pp.</p>
<p>Felix, L.P. & Guerra, M. (2010) Variation in chromosome number and the basic number of subfamily Epidendroideae (Orchidaceae). <em>Botanical Journal of the Linnaean Society</em> 163 (2): 234–278. https://doi.org/10.1111/j.1095-8339.2010.01059.x</p>
<p>Guerra, M. (1986) Reviewing the chromosome nomenclature of Levan <em>et al.</em> <em>Revista Brasileira de Genética</em> 9: 741–743.</p>
<p>Hágsater, E. (1984) Towards an Understanding of the Genus <em>Epidendrum</em>. <em>In:</em> Tan, K.W. (Ed.) <em>Proceedings of the Eleventh World Orchid Conference, Miami, U.S.A.</em> pp. 195–201.</p>
<p>Hágsater, E. & Krahl, A.H. (2020) <em>Epidendrum deditae</em>, a new species of Orchidaceae (Laeliinae) from northern Brazil. <em>Phytotaxa </em>440 (3): 225–231. https://doi.org/10.11646/phytotaxa.440.3.4</p>
<p>Hágsater, E. & Salazar, G.A. (1993) The genus<em> Epidendrum, </em>Part 1. A century of new species in<em> Epidendrum</em>. <em>Icones Orchidacearum 2</em>: pl. 101–200.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2018) <em>Epidendrum plurifolionocturnum</em>. <em>In:</em> Hágsater, E. & Santiago, E. (Eds.) The genus <em>Epidendrum</em>, Part 13. <em>Icones Orchidacearum</em> 16 (2): pl. 1695.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2001) The genus <em>Epidendrum</em>, Part 3. A third century of new species in <em>Epidendrum</em>.<em> Icones Orchidacearum </em>4: pl. 401–500.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2004) The genus <em>Epidendrum</em>, Part 4. A fourth century of new species in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>7: pl. 701–800.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2006) The Genus<em> Epidendrum</em>, Part 5. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>8: pl. 801–900.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2007) The genus <em>Epidendrum</em>, Part 6. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>9: pl. 901–1000.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2008) The genus <em>Epidendrum</em>, Part 7. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>11: pl. 1101–1200.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2009) The genus <em>Epidendrum</em>, Part 8. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>12: pl. 1201–1300.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2010) The genus <em>Epidendrum</em>, Part 9. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>13: pl. 1301–1400.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2013) The genus <em>Epidendrum</em>, Part 10. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>14: pl. 1401–1500.</p>
<p>Hágsater, E. & Sánchez-Saldaña, L. (2015) The Genus <em>Epidendrum</em>, Part 11. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>15 (1): pl. 1501–1568.</p>
<p>Hágsater, E. & Santiago, E. (2018a) The genus <em>Epidendrum</em>, Part 12. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>16 (1): pl. 1601–1667.</p>
<p>Hágsater, E. & Santiago, E. (2018b) The genus <em>Epidendrum</em>, Part 12. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum</em> 16 (2): pl. 1668–1700.</p>
<p>Hágsater, E. & Santiago, E. (2019) The genus <em>Epidendrum</em>, Part 12. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>17 (1): pl. 1701–1756.</p>
<p>Hágsater, E. & Santiago, E. (2020a) The genus <em>Epidendrum</em>, Part 13. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>17 (2): pl. 1757–1800.</p>
<p>Hágsater, E. & Santiago, E. (2020b) The genus <em>Epidendrum</em>, Part 14. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>18 (1): pl. 1849–1848.</p>
<p>Hágsater, E. & Santiago, E. (2021) The genus <em>Epidendrum</em>, Part 14. Species new & old in <em>Epidendrum</em>. <em>Icones Orchidacearum </em>18 (2): pl. 1801–1900.</p>
<p>Hágsater, E. & Soto-Arenas, M. (2005a) <em>Epidendrum</em>. <em>In</em>: Pridgeon, A.M., Cribb, P.J., Chase, M.W. & Rasmussen, F.N. (Eds.) <em>Genera Orchidacearum 4: Epidendroideae (Part 1)</em>. Oxford University Press, Oxford, pp. 236–251.</p>
<p>Hágsater, E. & Soto-Arenas, M. (2005b) Towards an understanding of diversification of the genus <em>Epidendrum. In: </em>Raynal-Roques, A., Roquenant, A. & Prat, D. (Eds.) <em>Proceedings of the 18th World Orchid Conference</em>, Dijon France, Naturalia, Abstract. pp. 225.</p>
<p>Hágsater, E., Archila, F. & Chiron, G. (2015) <em>Epidendrum ixilum</em>. <em>In:</em> Hágsater, E. & Santiago, E. (Eds.) The Genus <em>Epidendrum</em>, Part. 11. <em>Icones Orchidacearum </em>15 (1): pl. 1529.</p>
<p>Hágsater, E., Sánchez, L.S. & García-Cruz, J. (1999) The genus <em>Epidendrum</em>, Part 2. A second century of new species in <em>Epidendrum</em>. <em>Icones Orchidacearum</em> 3: pl. 301–400.</p>
<p>Lindley, J. (1834) <em>Epidendrum longicolle</em>. <em>Edwards’s Botanical Register </em>24 (Misc. 24): 49.</p>
<p>Linnaeus, C. (1759) <em>Epidendrum ciliare</em>. <em>Systema Naturae, Editio Decima </em>2: 1246.</p>
<p>Linnaeus, C. (1763) <em>Species Plantarum, ed. 2, vol. 2</em>. Imprensis Direct. Laurentii Salvii, Holmiae, 899 pp.</p>
<p>Moosburg, M., Hágsater, E. & Ochse, M. (2014) Ökologische Zusammenhänge anhand von Sammlungsexemplaren erkennen: Adhemarius sexoculata Grote, 1867, als Bestäuber bei der Orchideengattung <em>Epidendrum</em> Linnaeus, 1753 (Lepidoptera: Sphingidae). <em>Entomologische Zeitschrift, Stuttgart</em> 124: 2013–2014.</p>
<p>Mori, S.A., Silva, L.A., Lisboa, G. & Coradin, L. (1989) <em>Manual de Manejo do Herbário Fanerogâmico</em>. Ceplac, Ilhéus, 104 pp.</p>
<p>Pessoa, E.M. (2021) <em>Epidendrum</em>. <em>In:</em> Flora do Brasil 2020, Jardim Botânico do Rio de Janeiro, Brasil. Available from: http://floradobrasil.jbrj.gov.br/ (accessed 20 April 2021)</p>
<p>Pessoa, E.M., Cordeiro, J.M.P., Felix, L.P., Almeida, E.M., Costa, L., Nepomuceno, Á., Souza, G., Chase, M., Alves, M. & Van den Berg, C. (2020) Too many species: morphometrics, molecular phylogenetics and genome structure of a Brazilian species complex in <em>Epidendrum</em> (Laeliinae; Orchidaceae) reveal fewer species than previously thought. <em>Botanical Journal of the Linnaean Society</em> 195 (2): 161–188. https://doi.org/10.1093/botlinnean/boaa071</p>
<p>Salazar, G.A. & Dressler, R.L. (2011) The leaves got it right again: DNA phylogenetics supports a sister-group relationship between <em>Eurystyles </em>and<em> Lankesterella </em>(Orchidaceae: Spiranthinae). <em>Lankesteriana </em>11 (3): 337–347. https://doi.org/10.15517/lank.v11i3.18289</p>
<p>Sambin, A. & Aucourd, M. (2021) Mise à jour de l’inventaire des orchidées présentes autour et sur le pic Coudreau des Monts Bakra. Avec une note taxinomique pour quelques espèces. <em>Richardiana N.S. </em>5: 121–141.</p>
<p>Sambin, A. & Essers, D. (2018) Un nouvel <em>Epidendrum</em> (Orchidaceae) du groupe «Nocturnum» de Guyane. <em>Richardiana N.S. </em>2: 144–151.</p>
<p>Schneider, C.A., Rasband, W.S. & Eliceiri, K.W. (2012) NIH Image to ImageJ: 25 years of image analysis. <em>Nature Methods </em>9: 671–675. https://doi.org/10.1038/nmeth.2089</p>
<p>Schweinfurth, C. (1959) Orchids of Peru. <em>Fieldiana Botany</em> 30: 261–531.</p>