Skip to main content Skip to main navigation menu Skip to site footer
Type: Monograph
Published: 2022-06-17
Page range: 1-82
Abstract views: 70
PDF downloaded: 2

Thalloid Liverworts (Marchantiopsida) of Sri Lanka

Department of Botany, Faculty of Science, University of Peradeniya, Peradeniya, Sri Lanka, 20400; Postgraduate Institute of Science, University of Peradeniya, Peradeniya, Sri Lanka, 20400.
Department of Botany, Faculty of Science, University of Peradeniya, Peradeniya, Sri Lanka, 20400; Postgraduate Institute of Science, University of Peradeniya, Peradeniya, Sri Lanka, 20400.
Sri Lanka bryophytes complex thalloids simple thalloids

Abstract

A comprehensive taxonomic treatment of the thalloid liverworts of Sri Lanka is presented based on specimens collected from throughout the country covering all possible geographical localities. Eight families, nine genera, and seventeen species of complex thalloid liverworts and four families, six genera and seven species of simple thalloid liverworts are described from Sri Lanka. Taxonomic keys, descriptions, along with notes on ecology and distribution, and taxonomic illustrations are provided for all the taxa recognized. Plagiochasma appendiculatum Lehm. & Lindenb., Riccia crispatula Mitt., and Fossombronia cristula Austin, are reported as new to Sri Lanka.

References

<p>Abeywickrama, B.A. (1959) The genera of the liverworts of Ceylon. <em>Ceylon Journal of Science (Biological Sciences) </em>2: 33–81.</p>
<p>Abeywickrama, B.A. (1960) The genera of the mosses of Ceylon. <em>Ceylon Journal of Science (Biological Sciences) </em>3: 41–122.</p>
<p>Abeywickrama, B.A. &amp; Jansen, M.A.B. (1978a) A checklist of the liverworts of Sri Lanka. <em>UNESCO: Man and the Biosphere National Committee for Sri Lanka Publication </em>1: 1–10.</p>
<p>Abeywickrama, B.A. &amp; Jansen, M.A.B. (1978b) A checklist of the mosses of Sri Lanka. <em>UNESCO</em>: <em>Man and the Biosphere National Committee for Sri Lanka </em>2: 1–25.</p>
<p>Ah-Peng, C. &amp; Bardat, J. (2005) Check list of the bryophytes of Réunion Island (France). <em>Tropical bryology </em>26: 89–118.&nbsp; https://doi.org/10.11646/bde.26.1.14</p>
<p>Alam, A. (2014) Morphotaxonomy of three rare terricolous taxa of Jungermanniales occurring in Nilgiri hills (Western Ghats) India. <em>International Journal of Environment </em>3 (2): 263–275.&nbsp; https://doi.org/10.3126/ije.v3i2.10640</p>
<p>Alam, A. &amp; Srivastava, S.C. (2009) Current status of genus <em>Plagiochasma </em>in Nilgiri and Palni hills with semi details of spores. <em>Indian Journal of Forestry </em>32 (4): 623–634.</p>
<p>Alam, A., Vats, S. &amp; Behera, K.K. (2012) <em>Exormotheca ceylonensis </em>Meijer-a threatened liverwort in India, rediscovered in Palni Hills, Tamil Nadu. <em>Journal of threatened taxa </em>4 (5): 2593–2595.&nbsp; https://doi.org/10.11609/JoTT.o2611.2593-5</p>
<p>Allen, N.S &amp; Korpelainen, H. (2006) Notes on Neotropical <em>Cyathodium</em>. <em>Cryptogamie. Bryologie</em> 27 (1): 85–96.</p>
<p>Allen, N.S. (2001) <em>Cyathodium bischlerianum</em>, sp. nov. (Marchantiales), a new species from the Neotropics. <em>The Bryologist</em> 104 (1): 141–145.&nbsp; https://doi.org/10.1639/0007-2745(2001)104[0141:CBSNMA]2.0.CO;2</p>
<p>Allen, N.S., Lépiz, E. &amp; De Gracia, J.E. (2004) <em>Cyathodium foetidissimum </em>(Marchantiales), an Asiatic species new to Tropical America. <em>The Bryologist </em>107 (1): 41–46.&nbsp; https://doi.org/10.1639/0007-2745(2004)107[41:CFMAAS]2.0.CO;2</p>
<p>Andriessen, L., Sotiaux, A., Nagels, C. &amp; Sotiaux, O. (1995) <em>Aneura maxima </em>(Schiffn.) Steph. in Belgium, new for the European liverwort flora. <em>Journal of bryology </em>18 (4): 803–806.&nbsp; https://doi.org/10.1179/jbr.1995.18.4.803</p>
<p>Arzeni, C.B. (1965) Bryophytes of the cedar groves of Lebanon. <em>Bryologist </em>68 (1): 109–113.&nbsp; https://doi.org/10.2307/3240995</p>
<p>Ashton, P.M.S., Gamage, S., Gunatilleke, I.A.U.N. &amp; Gunatilleke, C.V.S. (1997) Restoration of a Sri Lankan rainforest: using Caribbean pine Pinus caribaea as a nurse for establishing late-successional tree species. <em>Journal of Applied Ecology</em> 34 (4): 915–925.&nbsp; https://doi.org/10.2307/2405282</p>
<p>Asthana, A.K. &amp; Sahu, V. (2013) Bryophyte Diversity in Mukteshwar (Uttarakhand): an overview. <em>Archive for Bryology </em>154: 1–11.</p>
<p>Austin, C.F. (1869) Characters of some new Hepaticae (mostly North American) together with, Notes on a new imperfectly described species. <em>Proceedings of the Academy of Natural Sciences of Philadelphia </em>21 (12): 228.&nbsp; https://doi.org/10.5962/bhl.title.40659</p>
<p>Bączkiewicz, A. &amp; Buczkowska, K. (2016) Differentiation and genetic variability of three cryptic species within the <em>Aneura pinguis</em> complex (Jungermanniidae, Marchantiophyta). <em>Cryptogamie, Bryologie</em> 37 (1): 3–18. https://doi.org/10.7872/cryb/v37.iss1.2016.3</p>
<p>Baczkiewicz, A., Sawicki, J., Buczkowska, K., Polok, K. &amp; Zielinski, R. (2008) Application of different DNA markers in studies on cryptic species of <em>Aneura pinguis </em>(Jungermanniopsida, Metzgeriales). <em>Cryptogamie Bryologie </em>29 (1): 3–21.</p>
<p>Buczkowska, K., Adamczak, M. &amp; Bączkiewicz, A. (2006) Morphological and anatomical differentiation within the <em>Aneura pinguis</em> complex (Metzgeriales, Hepaticae). <em>Biol Letter</em> 43 (1): 51–68.</p>
<p>Bag, A.K., Singh, S.K. &amp; Bhattacharya, S.G. (2007) Riccia of west Bengal. <em>Nelumbo</em> 49 (1–4): 173–186.</p>
<p>Bagwan, S.A. &amp; Kore, B.A. (2015) Species diversity of genus <em>Riccia</em> (Mich.) L. in Satara district (Maharashtra) India. <em>Plant Science Today</em> 2 (4): 187–191.</p>
<p>Bakalin, V.A., Arikawa, T. &amp; Higuchi, M. (2013) A Collection of Hepatics from the Tottori Prefecture, Japan. <em>Bulletin of the National Museum of Nature and Science, Series B (Botany), Tokyo </em>39 (4): 165–172.&nbsp; https://doi.org/10.15298/arctoa.22.19</p>
<p>Bapna, K.R. &amp; Kachroo, P. (2000) <em>Hepaticology in India </em>I<em>. </em>Himanshu Publications. Udaipur.</p>
<p>Bateman, R.M., Crane, P.R., Di Michele, W.A., Kenrick, P.R., Rowe, N.P., Speck, T. &amp; Stein, W.E. (1998) Early evolution of land plants: phylogeny, physiology, and ecology of the primary terrestrial radiation. <em>Annual Review of Ecology and Systematics </em>29 (1): 263–292.&nbsp; https://doi.org/10.1146/annurev.ecolsys.29.1.263</p>
<p>Benson-Evans, K. &amp; Hughes, J.G. (1955) The Physiology of Sexual Reproduction in <em>Lunularia cruciata </em>(L.) Dum. <em>Transactions of the British Bryological Society </em>2 (4): 513–522.&nbsp; https://doi.org/10.1179/006813855804829922</p>
<p>Bhattacharyya, S.D. (2011) Threats and Conservation of Liverworts in India: An Overview. <em>Assam University Journal of Science and Technology </em>7 (1): 168–172.</p>
<p>Bischler, H. (1979) <em>Plagiochasma </em>Lehm. et Lindenb. IV. Les taxa américains. <em>Revue Bryologique et Lichénologique </em>45 (3): 255–333.</p>
<p>Bischler, H. (1984) <em>Marchantia </em>L. The New World species. <em>Bryophytorum Bibliotheca </em>26.</p>
<p>Bischler-Causse, H. (1989) <em>Marchantia </em>L. The Asiatic &amp; Oceanic taxa. <em>Bryophytorum Bibliotheca </em>38: 1–318.</p>
<p>Bischler, H. (2004) Liverworts of the Mediterranean. <em>Bryphytorum Bibliotheca </em>61: 1–252.</p>
<p>Bischler, H. &amp; Sergio, C. (1984) Un neotvpe pour <em>Plagiochasma rupestre </em>(J R. &amp; G. Forst.) Steph. <em>Cnptogamie. Bryologie Lichenologie </em>5: 173–174.</p>
<p>Bischler-Causse, H. &amp; Boisselier-Dubayle, M.C. (1991) Lectotypification of <em>Marchantia polymorpha </em>L. <em>Journal of Bryology </em>16 (3): 361–365.&nbsp; https://doi.org/10.1179/jbr.1991.16.3.361</p>
<p>Bischler-Causse H &amp; Piippo, S. (1991) Bryophyte flora of Huon Peninsula, Papua New Guinea. L. (Marchantiaceae, Hepaticae). <em>Annales Botanici Fennici </em>28: 277–301.</p>
<p>Bischler-Causse, H., Gradstein, S.R., Jovet-Ast, S., Long, D.G. &amp; Allen, N.S. (2005) Marchantiidae. <em>Flora Neotropica, Monograph </em>97: 1–267.</p>
<p>Bischler, H. &amp; Jovet-Ast, S. (1981) The biological significance of morphological characters in Marchantiales (Hepaticae). <em>Bryologist </em>84 (2): 208–215.&nbsp; https://doi.org/10.2307/3242822</p>
<p>Blockeel, T.L. &amp; Nieuwkoop, J.A. (2016) The bryophyte flora of Lesbos, Greece. <em>Herzogia </em>29 (1): 1–34.&nbsp; https://doi.org/10.13158/heia.29.1.2016.1</p>
<p>Boisselier-Dubayle, M.C., Lambourdiere, J. &amp; Bischler, H. (1998) Taxa delimitation in <em>Reboulia </em>investigated with morphological, cytological, and isozyme markers. The <em>Bryologist </em>101 (1): 61–69.&nbsp; https://doi.org/10.2307/3244074</p>
<p>Bonner, C.E.B. (1953) De Hepaticis II. An unpublished section of volume 6 of Stephani’s Species Hepaticarum: the genus Marchantia. <em>Candollea</em> 14: 102–112.</p>
<p>Brown, R.C. &amp; Lemmon, B.E. (2008) γ-Tubulin and microtubule organization during meiosis in the liverwort <em>Ricciocarpus natans </em>(Ricciaceae). <em>American journal of botany </em>95 (6): 664–671.&nbsp; https://doi.org/10.3732/ajb.2007388</p>
<p>Buck, W.R., Goffinet, B. &amp; Shaw, A.J (2000) Novel relationships in pleurocarpous mosses as revealed by cpDNA sequences. <em>The Bryologist </em>103 (4): 774–789.&nbsp; https://doi.org/10.1639/0007-2745(2000)103[0774:NRIPMAR]2.0.CO;2</p>
<p>Buczkowska, K. &amp; Baczkiewicz, A. (2006) <em>Aneura maxima–</em>a liverwort new to Poland. <em>Cryptogamie Bryologie </em>27 (4): 453–458.</p>
<p>Campbell, D.H. (1936) The relationships of the Hepaticae. <em>The Botanical Review </em>2 (2): 53–66.&nbsp; https://doi.org/10.1007/BF02869921</p>
<p>Campbell, E.O. (1965) <em>Lunularia </em>in New Zealand. <em>Tuatara </em>13 (1): 31–41.&nbsp; https://doi.org/10.1080/00480169.1965.33590</p>
<p>Campbell, E.O. (1988) <em>Fossombronia </em>species in New Zealand. <em>Tuatara </em>30: 12–22.</p>
<p>Capesius, I. &amp; Bopp, M. (1997) New classification of liverworts based on molecular and morphological data. <em>Plant Systematics and Evolution</em> 207 (1-2): 87–97.&nbsp; https://doi.org/10.1007/BF00985210</p>
<p>Carreon, H.G., Morales, N.E., Cabras, A.A. &amp; Medina, M.N.D. (2016) Preliminary list of bryophytes in Tagbaobo, Kaputian, Island Garden City of Samal, Philippines. <em>University of Mindanao International Multdisciplinary Research Journal </em>1 (1): 152–157.</p>
<p>Cargill, D.C., Neal, W.C., Sharma, I. &amp; Gueidan, C. (2016) A preliminary molecular phylogeny of the genus <em>Riccia</em> L.(Ricciaceae) in Australia. <em>Australian Systematic Botany</em> 29 (3): 197–217.&nbsp; https://doi.org/10.1071/SB16018</p>
<p>Cavers, F. (1910) The inter-relationships of the Bryophyta. III. Anacrogynous Jungermanniales. <em>New Phytologist </em>9 (6): 197–234.</p>
<p>Cavers, F. (1911) The Classification of the Bryophyta X. New Phytologist 10: 42.&nbsp; https://doi.org/10.1111/j.1469-8137.1911.tb04944.x</p>
<p>Chantanaorrapint, S. &amp; Sridith, K. (2014) The genus <em>Plagiochasma </em>(Aytoniaceae, Marchantiopsida) in Thailand. <em>Cryptogamie Bryologie</em> 35 (2): 127–132.&nbsp; https://doi.org/10.7872/cryb.v35.iss2.2014.127</p>
<p>Coker, W.C. (1903) Selected notes. II. Liverworts. <em>Botanical Gazette </em>36 (3): 225–230.&nbsp; https://doi.org/10.1086/328398</p>
<p>Costa, D.P. (2008) Metzgeriaceae. <em>Flora Neotropica </em>102: 1–169.</p>
<p>Crandall-Stotler, B. (1980) Morphogenetic Designs and a Theory of Bryophyte Origins and Divergence. <em>BioScience </em>30 (9): 580–585.&nbsp; https://doi.org/10.2307/1308107</p>
<p>Crandall-Stotler, B. (1986) Morphogenesis, developmental anatomy and bryophyte phylogenetics: contraindications of monophyly. <em>Journal of bryology </em>14 (1): 1–23. https://doi.org/10.1179/jbr.1986.14.1.1</p>
<p>Crandall-Stotler, B., Stotler, R.E. &amp; Long, D.G. (2009) Phylogeny and classification of the Marchantiophyta. <em>Edinburgh journal of botany </em>66 (01): 155–198.&nbsp; https://doi.org/10.1017/S0960428609005393</p>
<p>Crandall-Stotler, B.J. &amp; Gradstein, S.R. (2017) A new riverine species of the liverwort <em>Fossombronia </em>(Pelliales, Fossombroniaceae) from Ecuador. <em>Bryophyte diversity and evolution </em>39 (1): 94–101.&nbsp; https://doi.org/10.11646/bde.39.1.13</p>
<p>Crandall-Stotler, B.J., Forrest, L. &amp; Stotler, R.E. (2005) Evolutionary trends in the simple thalloid liverworts (Marchantiophyta, Jungermanniopsida subclass Metzgeriidae). <em>Taxon </em>54 (2): 299–316.</p>
<p>&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; https://doi.org/10.2307/25065359</p>
<p>Crandall-Stotler, B. &amp; Stotler, R. (2000) Morphology and classification of the Marchantiophyta. <em>In</em>: Shaw, A.J. &amp; Goffinet, B. (Eds.) <em>Bryophyte Biology</em>. Cambridge: Cambridge University Press. pp. 21–70.</p>
<p>&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; https://doi.org/10.1017/CBO9781139171304.003</p>
<p>Crandall-Stotler, B., Stotler, R.E. &amp; Long, D.G. (2009) Phylogeny and classification of the Marchantiophyta. <em>Edinburgh Journal of Botany</em> 66 (1): 155–198.</p>
<p>&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; https://doi.org/10.1017/S0960428609005393</p>
<p>Crane, P.R. (1985) Phylogenetic analysis of seed plants and the origin of angiosperms. <em>Annals of the Missouri Botanical Garden </em>72 (4): 716–793.&nbsp; https://doi.org/10.2307/2399221</p>
<p>Cros, R.M., Sáez, L. &amp; Rosselló, J.A. (2005) <em>Plagiochasma appendiculatu</em>m Lehm. &amp; Lindenb. (Marchantiales, Aytoniaceae), a species new to the European bryophyte flora. <em>Journal of bryology </em>27 (1): 3–6.&nbsp; https://doi.org/10.1179/174328205X40527</p>
<p>Crum, H.A. (1991) <em>Liverworts and hornworts of southern Michigan</em>. University of Michigan Herbarium, Ann Arbor, MI. U.S.A.</p>
<p>Daniels, A.E.D. (2010) Checklist of the bryophytes of Tamil Nadu, India. <em>Archieve for Bryology </em>65.</p>
<p>Daskon, C and Binns, T. (2010) Culture, tradition and sustainable rural livelihoods:</p>
<p>exploring the culture-development interface in Kandy, Sri Lanka. <em>Community Development Journal </em>45 (4): 494–517.</p>
<p>Davis, E.C. (2004) A molecular phylogeny of leafy liverworts (Jungermanniidae: Marchantiophyta). <em>Monographs in Systematic Botany from the Missouri Botanical Garden</em> 98: 61–86.&nbsp; https://doi.org/10.1016/j.ympev.2004.06.006</p>
<p>Dissanayake, C.B. (1991) The fluoride problem in the ground water of Sri Lanka—environmental management and health. <em>International Journal of Environmental Studies </em>38 (2–3): 137–155.&nbsp; https://doi.org/10.1080/00207239108710658</p>
<p>Deutsch, H. (1912) A study of <em>Targionia hypophylla</em>. <em>Botanical Gazette </em>53 (6): 492–503.&nbsp; https://doi.org/10.1086/330847</p>
<p>Djamali, M., Kürschner, H., Akhani, H., de Beaulieu, J.L., Amini, A., Andrieu-Ponel, V., Ponel, P. &amp; Stevens, L (2008) Palaeoecological significance of the spores of the liverwort <em>Riella </em>(Riellaceae) in a late Pleistocene long pollen record from the hypersaline Lake Urmia, NW Iran. <em>Review of palaeobotany and palynology </em>152 (1–2): 66–73.&nbsp; https://doi.org/10.1016/j.revpalbo.2008.04.004</p>
<p>Duckett, J.G. &amp; Ligrone, R. (2006) <em>Cyathodium </em>Kunze (Cyathodiaceae: Marchantiales), a tropical liverwort genus and family new to Europe, in southern Italy. <em>Journal of bryology </em>28 (2): 88–96.&nbsp; https://doi.org/10.1179/174328206X105353</p>
<p>Duff, R.J. &amp; Nickrent, D.L. (1999) Phylogenetic relationships of land plants using mitochondrial small-subunit <em>r</em>DNA sequences. <em>American Journal of Botany </em>86 (3): 372–386.</p>
<p>&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; https://doi.org/10.2307/2656759</p>
<p>Dumortier, B.-C. (1829) <em>Analyse des plantes.</em> Tournay, Paris, France.</p>
<p>Edwards, D., Duckett, J.G. &amp; Richardson, J.B. (1995) Hepatic characters in the earliest land plants. <em>Nature </em>374 (6523): 635–636.&nbsp; https://doi.org/10.1038/374635a0</p>
<p>Eriyagama, N., Smakhtin, V., Chandrapala, L. &amp; Fernando, K. (2010) <em>Impacts of climate change on water resources and agriculture in Sri Lanka: a review and preliminary vulnerability mapping, </em>IWMI, 135.</p>
<p>Evans, A.W. (1915) The genus <em>Plagiochasma </em>and its North American species. <em>Bulletin of the Torrey botanical club </em>42 (5): 259–308.&nbsp; https://doi.org/10.2307/2479766</p>
<p>Evans, A.W. (1917) Preliminary list of Arizona hepaticae. <em>The Bryologist </em>20 (4): 60–62.&nbsp; https://doi.org/10.2307/3238013</p>
<p>Evans, A.W. (1919) A taxonomic study of <em>Dumortiera</em>. <em>Bulletin of the Torrey Botanical Club </em>46 (5): 167–182.&nbsp; https://doi.org/10.2307/2480346</p>
<p>Farmer, J.B. (1894) Studies in Hepaticae: On <em>Pallavicinia decipiens</em> Mitten. <em>Annals of Botany</em> 8 (29): 35–52.&nbsp; https://doi.org/10.1093/oxfordjournals.aob.a090698</p>
<p>Fernando, C.H. (1986) <em>Ecology and Biogeography in Sri Lanka. </em>Dr. W. Junk Publishers,The Hague, The Netherlands.</p>
<p>Forrest, L.L. &amp; Crandall-Stotler, B.J. (2005) Progress towards a robust phylogeny for the liverworts, with particular focus on the simple thalloids (Selected Papers Presented at the 15 World Congress on Bryology Held at Merida, Venezuela, January 12–16, 2004). <em>Journal of the Hattori Botanical Laboratory </em>97: 127–159.</p>
<p>Forrest, L.L., Davis, E.C., Long, D.G., Crandall-Stotler, B.J., Clark, A. &amp; Hollingsworth, M.L. (2006) Unraveling the evolutionary history of the liverworts (Marchantiophyta): multiple taxa, genomes and analyses. <em>The bryologist</em> 109 (3): 303–334.&nbsp; https://doi.org/10.1639/0007-2745(2006)109[303:UTEHOT]2.0.CO;2</p>
<p>Forrest, L.L., Schuette, S.W., Crandall-Stotler, B.J. &amp; Stotler, R.E. (2005) A molecular study of the simple thalloid liverwort <em>Jensenia</em> (Marchantiophyta, Pallaviciniaceae). <em>The Bryologist</em> 108 (2): 204–211.&nbsp; https://doi.org/10.1639/0007-2745(2005)108[0204:AMSOTS]2.0.CO;2</p>
<p>Frey, W. &amp; Kürschner, H. (1983) Contributions towards a bryophyte flora of Iran. New records from Iran. <em>Iranian Journal Botany </em>2 (1): 13–19.</p>
<p>Frey, W. &amp; Kürschner, H. (2011) Asexual reproduction, habitat colonization and habitat maintenance in bryophytes. <em>Flora-Morphology, Distribution, Functional Ecology of Plants</em> 206 (3): 173–184. https://doi.org/10.1016/j.flora.2010.04.020</p>
<p>Frye, T. &amp; Clark, L. (1937) Hepaticae of North America.<em> University of Washington Publications in Biology</em> 6 (1–5): 1–1018.</p>
<p>Furuki, T. (2006) Taxonomical studies of the family Aneuraceae (Hepaticae) based on the Philippine collections made by Dr. and Mrs. AJ Sharp and Dr. Z. Iwatsuki. <em>The Journal of the Hattori Botanical Laboratory</em> 100: 89–99.</p>
<p>Furuki, T. &amp; Dalton, P.J. (2008) <em>Vandiemenia ratkowskiana</em> Hewson (Marchantiophyta): a revised description and reassessment of its taxonomic status. <em>Journal of Bryology</em> 30 (1): 48–54.&nbsp; https://doi.org/10.1179/174328208X282139</p>
<p>Furuki, T. &amp; Long, D.G. (1994) <em>Aneura crateriformis</em>, a new liverwort species from the East Himalaya and China. <em>Journal of Bryology</em> 18 (2): 281–286. https://doi.org/10.1179/jbr.1994.18.2.281</p>
<p>Ghahreman, A., Faridi, M., Shirzadian, S. &amp; Attar, F. (2007) New and interesting moss records for Iran. <em>Turkish Journal of Botany</em> 31 (1): 41–48.</p>
<p>Gledhill, D. (2008) <em>The names of plants</em>. Cambridge University Press.&nbsp; https://doi.org/10.1017/CBO9780511550898</p>
<p>Glime, J.M. (2015) Water relations: conducting structures. <em>In</em>: Glime, J.M. (ed.) <em>Bryophyte ecology. </em>Volume I. Ebook sponsored by Michigan Technological University and the International Association of Bryologists.</p>
<p>Goffinet, B., Buck, W.R. &amp; Shaw, A.J. (2009) Morphology and classification of the Bryophyta. <em>In</em>: Goffinet, B. &amp; Shaw, A.J. (Eds.) <em>Bryophyte Biology. </em>Cambridge University Press.&nbsp; https://doi.org/10.1017/CBO9780511754807</p>
<p>Gradstein, S.R. (2013) Afro-American hepatics revisited. <em>Polish botanical journal </em>58 (1): 149–177.&nbsp; https://doi.org/10.2478/pbj-2013-0016</p>
<p>Gradstein, S.R. &amp; Ziemmeck, F. (2014) CDF checklist of Galapagos horn-&amp; liverworts. <em>In</em>: Bungartz, F., Herrera, H., Jaramillo, P., Tirado, N., Jiménez-Uzcátegui, G., Ruiz, D., Guézou, A. &amp; Ziemmeck, F. (Eds.) <em>Charles Darwin Foundation Galapagos Species Checklist</em>—<em>Lista de Especies de Galápagos de la Fundación Charles Darwin. </em>Charles Darwin Foundation.</p>
<p>Gradstein, S.R. &amp; Cuvertino, J. (2015) Observations on the liverwort flora of the surroundings of Santiago, Central Chile. <em>Cryptogamie Bryologie </em>36 (2): 129–141.</p>
<p>&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; https://doi.org/10.7872/cryb.v36.iss2.2015.129</p>
<p>Gray, S.F. (1821) <em>A natural arrangement o f British plants </em>1. London. pp. 1–824.</p>
<p>Groth-Malonek, M. &amp; Knoop, V. (2005) Bryophytes and other basal land plants: the mitochondrial perspective. <em>Taxon </em>54 (2): 293–297.&nbsp; https://doi.org/10.2307/25065461</p>
<p>Gunatilleke, N., Pethiyagoda, R. &amp; Gunatilleke, S. (2008) Biodiversity of Sri Lanka. <em>Journal of the National Science Foundation of Sri Lanka </em>36 (25): 25–61.</p>
<p>Gunawardene, N.R., Daniels, D.A., Gunatilleke, I.A.U.N., Gunatilleke, C.V.S., Karunakaran, P.V., Nayak, G.K., Prasad, S., Puyravaud, P., Ramesh, B.R., Subramanian, K.A. &amp; Vasanthy, G. (2007) A brief overview of the Western Ghats–Sri Lanka. biodiversity hotspot. <em>Current science </em>93 (11): 1567–1572.</p>
<p>Hattori, S. (1968) Hepaticae collected by F. Schmid in Ceylon and Pakistan. <em>Candollea </em>23: 287–294.</p>
<p>Hassel de Menendez, G.G. (1963) Estudio de las Anthocerotales y Marchantiales de la Argentina. <em>Opera lilloana </em>7: 1–297.</p>
<p>Haupt, A.W. (1918) A morphological study of <em>Pallavicinia lyellii</em>. <em>Botanical Gazette </em>66 (6): 524–533.&nbsp; https://doi.org/10.1086/332375</p>
<p>Heinrichs, J., Gradstein, S.R., Wilson, R. &amp; Schneider, H. (2005) Towards a natural classification of liverworts (Marchantiophyta) based on the chloroplast gene <em>rbcL</em>. <em>Cryptogamie Bryologie </em>26: 131–150.</p>
<p>Heinrichs, J., Hentschel, J., Feldberg, K., Bombosch, A. &amp; Schneider, H. (2009) Phylogenetic biogeography and taxonomy of disjunctly distributed bryophytes. <em>Journal of Systematics and Evolution </em>47 (5): 497–508.&nbsp; https://doi.org/10.1111/j.1759-6831.2009.00028.x</p>
<p>Hernick, L.V., Landing, E. &amp; Bartowski, K.E. (2008) Earth’s oldest liverworts <em>Metzgeriothallus sharonae </em>sp. nov. from the Middle Devonian (Givetian) of eastern New York, USA. <em>Review of Palaeobotany and Palynology </em>148 (2): 154–162.&nbsp; https://doi.org/10.1016/j.revpalbo.2007.09.002</p>
<p>Hofmeister, W. (1851) <em>Vergleichende Untersuchungen der Keimung, Entfaltung und Fruchtbildung höherer Kryptogamen (Moose, Farrn, Equisetaceen, Rhizocarpeen und Lycopodiaceen) und die Samenbildung der Coniferen.</em> Leipzig</p>
<p>Hong, W.S. (2003) The Hepaticae and Anthocerotae of the Korean peninsula: identification keys to the taxa. <em>Lindbergia</em> 28: 134–147.</p>
<p>Kachroo, P. (1956) Morphology of <em>Pallavicinia </em>with reference to its species problem and the individuality of Pallaviciniaceae. <em>Proceedings of the National Academy of Sciences, India Section B: Biological Sciences </em>22: 6–21.</p>
<p>Katagiri, T. &amp; Mizutani, M. (2014) <em>Plagiochasma appendiculatum </em>Lehm. &amp; Lindenb., a species new to Japan. <em>Hattoria </em>5: 95–100.</p>
<p>Kenrick, P. &amp; Crane, P.R. (1997) The origin and early evolution of plants on land. <em>Nature </em>389 (6646): 33–39.&nbsp; https://doi.org/10.1038/37918</p>
<p>Khan, S.A. (1957) Studies in Ricciaceae of East Pakistan: I. New and Little Known Species of Riccia. <em>The Bryologist</em> 60 (1): 28–32.&nbsp; https://doi.org/10.2307/3240049</p>
<p>Knoop, V. (2004) The mitochondrial DNA of land plants: peculiarities in phylogenetic perspective. <em>Current genetics </em>46 (3): 123–139.&nbsp; https://doi.org/10.1007/s00294-004-0522-8</p>
<p>Kolkwitz, R. (1935) <em>Pflanzenphysiologie: Versuche und Beobachtungen an höheren und niederen Pflanzeneinschließlich Bakteriologie und Hydrobiologiemit Planktonkunde.</em> J. G. Fischer.</p>
<p>Krayesky, D.M., Crandall-Stotler, B. &amp; Stotler, R.E. (2005) A revision of the genus <em>Fossombronia</em> Raddi in East Asia and Oceania. <em>The Journal of the Hattori Botanical Laboratory</em> 98: 1–45.</p>
<p>Kreier, H., Feldberg, K., Mahr, F., Bombosch, A., Alexander, R. Schmidt, Rui-Liang Zhu, von Konrat, M., Shaw, B., Shaw, A.J. &amp; Heinrichs, J. (2010) Phylogeny of the leafy liverwort <em>Ptilidium</em>: cryptic speciation and shared haplotypes between the Northern and Southern Hemispheres. <em>Molecular Phylogenetics and Evolution </em>57 (3): 1260–1267.&nbsp; https://doi.org/10.1016/j.ympev.2010.10.002</p>
<p>Kugita, M., Kaneko, A., Yamamoto, Y., Takeya, Y., Matsumoto, T. &amp; Yoshinaga, K. (2003) The complete nucleotide sequence of the hornwort (<em>Anthoceros</em> <em>formosae</em>) chloroplast genome: insight into the earliest land plants. <em>Nucleic Acids Research </em>31 (2): 716–721.&nbsp; https://doi.org/10.1093/nar/gkg155</p>
<p>Kürschner, H. (2000) Bryophyte flora of the Arabian Peninsula and Socotra. <em>Nova Hedwigia </em>71 (1–2): 267.&nbsp; https://doi.org/10.1127/nova.hedwigia/71/2001/253</p>
<p>Kürschner, H. (2003) Further new bryophytes to Socotra and the Yemen mainland, including <em>Papillaria crocea </em>(Hampe) A. Jaeger (Meteoriaceae, Bryopsida), a palaeotropical Asian species. <em>Nova Hedwigia </em>76 (1–2): 257–267.&nbsp; https://doi.org/10.1127/0029-5035/2003/0076-0257</p>
<p>Kürschner, H. (2004) Life strategies and adaptations in bryophytes from the Near and Middle East. <em>Turkish Journal of Botany </em>28 (1–2): 73–84.</p>
<p>Kürschner, H. (2006) A bryophyte flora of Socotra Island, Yemen. <em>Englera </em>28: 97–162.&nbsp; https://doi.org/10.2307/20358185</p>
<p>Kuwahara, Y. (1978) Reduction of the female gametophyte in <em>Metzgeria decipiens</em> (Massal.) Schiffn. &amp; Gotts. and <em>M. foliicola</em> Schiffn.(Hepaticae). <em>Bulletin of the Torrey Botanical Club</em> 130–133.&nbsp; https://doi.org/10.2307/2484430</p>
<p>Kuwahara, Y. (1986) The Metzgeriaceae of the Neotropics. <em>Bryophytorum Bibliotheca</em> 28: 1–254.</p>
<p>Leitgeb, H. (1874) <em>Untersuchungen über die Lebermoose</em>, I–V. Jena: O. Deistungs Buchhandlung. https://doi.org/10.5962/bhl.title.10206</p>
<p>Lewis, L.A., Mishler, B.D. &amp; Vilgalys, R. (1997) Phylogenetic relationships of the liverworts (Hepaticae), a basal embryophyte lineage, inferred from nucleotide sequence data of the chloroplast gene <em>rbc</em>L. <em>Molecular phylogenetics and evolution </em>7 (3): 377–393.&nbsp; https://doi.org/10.1006/mpev.1996.0395</p>
<p>Li, R., Sun, B., Wang, H., He, Y., Yang, G., Yan, D. &amp; Lin, Z. (2014) <em>Marchantites huolinhensis </em>sp. nov. (Marchantiales)-A new fossil liverwort with gemma cups from the Lower Cretaceous of Inner Mongolia, China. <em>Cretaceous Research </em>50: 16–26.&nbsp; https://doi.org/10.1016/j.cretres.2014.03.007</p>
<p>Linnaeus, C. (1753) <em>Species Plantarum </em>ed.1. Laurentii Salvii, Holmiae [Stockholm] 1: 561–1200.</p>
<p>Little, E.L. (1949) Nomina conservanda proposals in Hepaticae. <em>The Bryologist</em> 52 (1): 1–22.&nbsp; https://doi.org/10.2307/3239727</p>
<p>Long, D.G. (2006) New higher taxa of complex thalloid liverworts (Marchantiophyta–Marchantiopsida). <em>Edinburgh Journal of Botany </em>63 (2–3): 257–262.&nbsp; https://doi.org/10.1017/S0960428606000606</p>
<p>Long, D.G., Forrest, L.L., Villarreal, J.C. &amp; Crandall-Stotler, B.J. (2016) Taxonomic changes in Marchantiaceae, Corsiniaceae and Cleveaceae (Marchantiidae, Marchantiophyta). <em>Phytotaxa </em>252 (1): 77–80.&nbsp; https://doi.org/10.11646/phytotaxa.252.1.9</p>
<p>Long, D. &amp; Rubasinghe, S.C.K. (2014) Liverworts and Hornworts of Sri Lanka: a revised checklist. <em>Ceylon Journal of Science (Biological Sciences) </em>43 (1): 1–36.&nbsp; https://doi.org/10.4038/cjsbs.v43i1.7280</p>
<p>Lotsy, J.P. (1910) Phylogeny of Plants. <em>Botanical Gazette </em>49 (6): 460–461.&nbsp; https://doi.org/10.1086/330244</p>
<p>Mallick, P. (2015) New Record of the four species of <em>Riccia</em> from Bankura district, West Bengal (India). <em>Plant Science Today</em> 2 (4): 175–178.&nbsp; https://doi.org/10.14719/pst.2015.2.4.142</p>
<p>Mamontov, Y.S., Konstantinova, N.A., Vilnet, A.A. &amp; Bakalin, V.A. (2015) On the phylogeny and taxonomy of Pallaviciniales (Marchantiophyta), with overview of Russian species. <em>Arctoa </em>24 (1): 98–123.&nbsp; https://doi.org/10.15298/arctoa.24.12</p>
<p>Meagher, D. (2008) Chapter Twenty-one: An Etymology of Australian Bryophyte Genera. 1–Liverworts and Hornworts. <em>Fieldiana Botany </em>45: 257–269.&nbsp; https://doi.org/10.3158/0015-0746-47.1.257</p>
<p>Meegaskumbura, M., Bossuyt, F., Pethiyagoda, R., Manamendra-Arachchi, K., Bahir, M., Milinkovitch, M.C. &amp; Schneider, C.J. (2002) Sri Lanka: an amphibian hot spot. <em>Science </em>298 (5592): 379–379.&nbsp; https://doi.org/10.1126/science.298.5592.379</p>
<p>Mehra, P.N. (1957) A New Suggestion on the Origin of Thallus in the Marchantiales. II. The Theory. <em>American Journal of Botany </em>44 (7): 573–581.&nbsp; https://doi.org/10.1002/j.1537-2197.1957.tb10578.x</p>
<p>Mehra, P.N. &amp; Vashisht, B.R. (1950) Embryology of <em>Petalophyllum indicum</em> Kash. and a new suggestion of the evolution ofthalloid habit from foliose forms. <em>The Bryologist</em> 53: 89–114. https://doi.org/10.1639/0007-2745(1950)53[90:EOPIKA]2.0.CO;2</p>
<p>Meijer, W. (1956) A new species of <em>Exormotheca </em>from Ceylon. <em>Journal of the Hattori Botanical Laboratory </em>16: 72–74.</p>
<p>Miller, H.A. &amp; Inoue, H. (1966) <em>Jensenia macrogyna</em>, a New Philippine Hepatic. <em>American Journal of Botany </em>53 (7): 708–711.&nbsp; https://doi.org/10.1002/j.1537-2197.1966.tb14027.x</p>
<p>Mishler, B.D. (2001) The biology of bryophytes—bryophytes aren’t just small tracheophytes. <em>American Journal of Botany </em>88 (11): 2129–2131.&nbsp; https://doi.org/10.2307/3558438</p>
<p>Mishler, B.D. &amp; Churchill, S.P. (1984) A cladistic approach to the phylogeny of the “bryophytes”. <em>Brittonia </em>36 (4): 406–424.&nbsp; https://doi.org/10.2307/2806602</p>
<p>Mishler, B.D. &amp; Churchill, S.P. (1985) Transition to a land flora: phylogenetic relationships of the green algae and bryophytes. <em>Cladistics </em>1 (4): 305–328.&nbsp; https://doi.org/10.1111/j.1096-0031.1985.tb00431.x</p>
<p>Mitten, W. (1861) Hepaticae Indiae Orientalis: an Enumeration of the Hepaticae of the East Indies.(continued). <em>Botanical Journal of the Linnean Society</em> 5 (19): 109–128.&nbsp; https://doi.org/10.1111/j.1095-8312.1861.tb01343.x</p>
<p>Miyoshi, N. (1966) Spore morphology of Hepaticae in Japan. <em>Bull. Okayama Coll. Sci. </em>2: 146.</p>
<p>Myers, N. (1990) The biodiversity challenge: expanded hot-spots analysis. <em>Environmentalist </em>10 (4): 243–256.&nbsp; https://doi.org/10.1007/BF02239720</p>
<p>Na-Thalang, O. (1980) A revision of the genus <em>Riccia</em> (Hepaticae) in Australia<em>. Brunonia</em> 3 (1): 61–140.&nbsp; https://doi.org/10.1071/BRU9800061</p>
<p>Nickrent, D.L., Parkinson, C.L., Palmer, J.D. &amp; Duff, R.J. (2000) Multigene phylogeny of land plants with special reference to bryophytes and the earliest land plants. <em>Molecular Biology and Evolution </em>17 (12): 1885–1895.&nbsp; https://doi.org/10.1093/oxfordjournals.molbev.a026290</p>
<p>Nunik, S.A. &amp; Gradstein, S.R. (2007) Wallace’s line and the distribution of the liverworts of Sulawesi. <em>Cryptogamie, Bryologie </em>28 (1): 3–14.</p>
<p>O’Shea, B. (2002) Checklist of the mosses of Sri Lanka. <em>Journal of Hattori Botanical Laboratory </em>92: 125–164.</p>
<p>O’Shea, B.J. (2003) Biogoegraphical relationships of the mosses of Sri Lanka. <em>Journal of Hattori Botanical Laboratory </em>93: 293–304.</p>
<p>O’Shea, B.J. &amp; Buck, W.R. (2001) Bryophytes of Uganda. 5. Bryocrumia LE Anderson (Hypnaceae), a monotypic moss genus new to Africa. <em>Tropical bryology </em>20: 103–108.&nbsp; https://doi.org/10.11646/bde.20.1.18</p>
<p>Onraedt, M. (1981) Bryophytes recoltees a Sri Lanka (Ceylan) V. <em>Journal of the Hattori botanical laboratory </em>50: 191–216.</p>
<p>Pande, S.K. &amp; Udar, R. (1958) Genus <em>Riccia</em> in India-II. Species of <em>Riccia</em> from South India with description of a new species and notes on the synonyms of some recently described ones. <em>Proceedings of the National Institute of Science of India, part B, Biological Sciences</em> 24 (2): 79–88.</p>
<p>Paton, J.A. (1999) <em>Liverwort flora of the British Isles</em>. Harley Books.</p>
<p>Perold, S.M. (1993) Studies in the Marchantiales (Hepaticae) from Southern Africa. 1. The genus <em>Dumortiera </em>and <em>D. hirsuta</em>; the genus <em>Lunularia </em>and <em>L. cruciata</em>. <em>Bothalia </em>23 (1): 49–57.&nbsp; https://doi.org/10.4102/abc.v23i1.783</p>
<p>Perold, S.M. (1995) Studies in the Marchantiales (Hepaticae) from southern Africa. 9, The genus <em>Marchantia </em>and its five local species. <em>Bothalia </em>25 (2): 183–204.&nbsp; https://doi.org/10.4102/abc.v25i2.727</p>
<p>Perold, S.M. (2001) Studies in the liverwort family Aneuraceae (Metzgeriales) from southern Africa. 1. The genus <em>Aneura </em>and its local representative. <em>Bothalia </em>31 (2): 167–173.&nbsp; https://doi.org/10.4102/abc.v31i2.515</p>
<p>Phephu, N. &amp; Rooy, J.V. (2013) Studies in the liverwort family Metzgeriaceae (Metzgeriales) from southern Africa. 1. <em>Metzgeria nudifrons </em>Stephani and a key to the species. <em>Polish Botanical Journal </em>58 (2): 449–455.&nbsp; https://doi.org/10.2478/pbj-2013-0045</p>
<p>Piippo, S. (1991) Bryophyte flora of the Huon Peninsula, Papua New Guinea. XXXIX. <em>Fossombronia </em>(Fossombroniaceae) and Metzgeria (Metzgeriaceae, Hepaticae). <em>Acta Botanica Fennica </em>143: 1–22.</p>
<p>Piippo, S., He, X.L. &amp; Koponen, T. (1997) Hepatics from northwestern Sichuan, China, with a checklist of Sichuan hepatics. <em>Annales Botanici Fennici</em>: 51–63.</p>
<p>Pradhan, N. &amp; Joshi, S.D. (2010) Liverworts and hornworts of Nepal: a synopsis. <em>Botanica Orientalis</em>–<em>Journal of Plant Science </em>6: 69–75.&nbsp; https://doi.org/10.3126/botor.v6i0.2913</p>
<p>Preußing, M., Olsson, S., Schäfer-Verwimp, A., Wickett, N.J., Wicke, S., Quandt, D. &amp; Nebel, M. (2010) New insights in the evolution of the liverwort family Aneuraceae (Metzgeriales, Marchantiophyta), with emphasis on the genus <em>Lobatiriccardia</em>. <em>Taxon</em> 59 (5): 1424–1440. https://doi.org/10.1002/tax.595009</p>
<p>Qiu, Y.L., Cho, Y., Cox, J.C. &amp; Palmer, J.D. (1998) The gain of three mitochondrial introns identifies liverworts as the earliest land plants. <em>Nature </em>394 (6694): 671–674.&nbsp; https://doi.org/10.1038/29286</p>
<p>Qiu, Y.L. &amp; Palmer, J.D. (1999) Phylogeny of early land plants: insights from genes and genomes. <em>Trends in plant Science </em>4 (1): 26–30.&nbsp; https://doi.org/10.1016/S1360-1385(98)01361-2</p>
<p>Qiu, Y., Li, L., Wang, B., Chen, Z., Knoop, V., Groth-Malonek, M., Dombrovska, O., Lee, J., Kent, L., Rest, J., Estabrook, G.F., Hendry, T.A., Taylor, D.W., Testa, C.M., Ambros, M., Crandall-Stotler, B., Duff, R.J., Stech, M., Frey, W., Quandt, D. &amp; Davis, C.C. (2006) The deepest divergences in land plants inferred from phylogenomic evidence. <em>Proceedings of the National Academy of Science</em> 103 (42): 15511−15516.</p>
<p>&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; https://doi.org/10.1073/pnas.0603335103</p>
<p>Qiu, Y.L., Li, L., Wang, B., Chen, Z., Dombrovska, O., Lee, J., Kent, L., Li, R., Jobson, R.W., Hendry, T.A., Taylor, D.W., Testa, C.M. &amp; Ambros, M. (2007) A nonflowering land plant phylogeny inferred from nucleotide sequences of seven chloroplast, mitochondrial, and nuclear genes. <em>International Journal of Plant Sciences </em>168 (5): 691–708.&nbsp; https://doi.org/10.1086/513474</p>
<p>Rabeau, L., Gradstein, S.R., Dubuisson, J.Y., Nebel, M., Quandt, D. &amp; Reeb, C. (2017) New insights into the phylogeny and relationships within the worldwide genus Riccardia (Aneuraceae, Marchantiophytina). <em>European Journal of Taxonomy </em>273: 1–26.&nbsp; https://doi.org/10.5852/ejt.2017.273</p>
<p>Ramaiya, M., Johnson, M.G., Shaw, B., Heinrichs, J., Hentschel, J., von Konrat, M., Davison, P.G. &amp; Shaw, A.J., (2010) Morphologically cryptic biological species within the liverwort <em>Frullania asagrayana</em>. <em>American journal of botany </em>97 (10): 1707–1718.&nbsp; https://doi.org/10.3732/ajb.1000171</p>
<p>Reeb, C. &amp; Bardat, J. (2014) Studies on African Riccardia types and related material.&nbsp;<em>Cryptogamie, Bryologie</em>&nbsp;35 (1): 47–75. https://doi.org/10.7872/cryb.v35.iss1.2014.47</p>
<p>Renzaglia, K.S. (1982) A comparative developmental investigation of the gametophyte generation in the Metzgeriales (Hepatophyta). <em>Bryophytorum Bibliotheca </em>24: 1–253. [J. Cramer. Vaduz.]</p>
<p>Renzaglia, K.S. &amp; Bartholomew, S.E. (1985) Sporeling development in <em>Fossombronia cristula </em>Aust. with special reference to the apical organization and growth. <em>The Bryologist </em>88 (4): 337–343.&nbsp; https://doi.org/10.2307/3242669</p>
<p>Renzaglia, K.S., Duff, R.J., Nickrent, D.L. &amp; Garbary, D.J. (2000) Vegetative and reproductive innovations of early land plants: implications for a unified phylogeny. <em>Philosophical Transactions of the Royal Society of London B: Biological Sciences </em>355 (1398): 769–793.&nbsp; https://doi.org/10.1098/rstb.2000.0615</p>
<p>Rubasinghe, S.C.K. (2011) Phylogeny and Taxonomy of the complex thalloid liverwort family Cleveaceae Cavers. Thesis (PhD). University of Edinburgh, Scotland, U.K.</p>
<p>Rubasinghe, S.C. &amp; Long, D.G. (2014) Bryophytes of Sri Lanka: a review of past exploration and taxonomic research and priorities for the future. <em>Journal of Bryology </em>36 (4): 259–270.&nbsp; https://doi.org/10.1179/1743282014Y.0000000121</p>
<p>Rubinstein, C.V., Gerrienne, P., de la Puente, G.S.L., Astini, R.A. &amp; Steemans, P. (2010) Early Middle Ordovician evidence for land plants in Argentina (eastern Gondwana). <em>New Phytologist </em>188 (2): 365–369.&nbsp; https://doi.org/10.1111/j.1469-8137.2010.03433.x</p>
<p>Ruklani, N.C.S. &amp; Rubasinghe, S.C.K. (2013) A Preliminary Survey of Bryophytes in the Central Province of Sri Lanka. <em>Ceylon Journal of Science (Biological Sciences) </em>42 (1): 67–72.&nbsp; https://doi.org/10.4038/cjsbs.v42i1.5901</p>
<p>Ruklani, N.C.S., Rubasinghe, S.C.K. &amp; Long, D.G. (2015) Morphological diversity of complex thalloid liverwort genera of Sri Lanka. <em>Ceylon Journal of Science (Biological. Sciences) </em>44 (2): 27–44.&nbsp; https://doi.org/10.4038/cjsbs.v44i2.7348</p>
<p>Ruklani, N.C.S., Rubasinghe, S.C.K. &amp; Villarreal, J.C. (2016) Two new records of Sri Lankan hornworts, <em>Notothylas javanica </em>(Notothyladaceae) and <em>Megaceros flagellaris </em>(Dendrocerotaceae). <em>Cryptogamie, Bryologie </em>37 (4): 435–444.&nbsp; https://doi.org/10.7872/cryb/v37.iss4.2016.435</p>
<p>Sahu, V. &amp; Asthana, A.K. (2015) Bryophyte diversity in Terai regions of Uttar Pradesh, India with some new additions to the state. <em>Tropical Plant Research</em> 2 (3): 180–191.</p>
<p>Santarelli, E. (1958) Contributo Alla Flora Crittogamica Versiliese: Epatiche. <em>Webbia </em>14 (1): 53–71.&nbsp; https://doi.org/10.1080/00837792.1958.10669684</p>
<p>Santos, N.D.D. &amp; Costa, D.P.D. (2010) Phytogeography of the liverwort flora of the Atlantic Forest of Southeastern Brazil. <em>Journal of Bryology </em>32 (1): 9–22.&nbsp; https://doi.org/10.1179/037366810X12578498135634</p>
<p>Schaumann, F., Frey, W., Pfeiffer, T. &amp; Stech, M. (2005) Molecular circumscription, intrafamilial relationships and biogeography of the Gondwanan liverwort family Pallaviciniaceae (Hepaticophytina, Bryophyta) Studies in austral temperate rain forest bryophytes 27. <em>Plant Systematics and Evolution </em>252 (1–2): 27–48.&nbsp; https://doi.org/10.1007/s00606-004-0279-z</p>
<p>Schuette, S.W. &amp; Stotler, R.E. (2005) A conspectus of the liverwort genus <em>Jensenia</em> in Latin America. <em>The Journal of the Hattori Botanical Laboratory</em> 97: 299–308.</p>
<p>Schiffner, V. (1893) Jungermanniaceae Akroginae. <em>In</em>: Engler, A. &amp; Prantl, K. (Eds.) <em>Die Nattirlichen Pflanzen-familien </em>1 (3). Wilhelm Engelmann, Leip-zig. pp. 61–134.</p>
<p>Schiffner, V., Pandé, S.K. &amp; Srivastava, K.P. (1959) The genus <em>Riccardia </em>Gray in India. II. <em>Riccardia palmatiformis </em>Schffn. and <em>R. decolyana </em>Schffn. <em>Journal of the Indian Botanical Society </em>38 (4): 538–542.</p>
<p>Schill, D.B. (2006) <em>Taxonomy and phylogeny of the liverwort genus Mannia (Marchantiophyta, Aytoniaceae). </em>PhD Thesis, University of Edinburgh, Edinburgh. [https://www.era.lib.ed.ac.uk/handle/1842/7463]</p>
<p>Schofield, W.B. (1985) <em>Introduction to bryology</em>. Macmillan Publishing Company. London.</p>
<p>Schuster, R.M. (1953) <em>Boreal Hepaticae: A manual of the liverworts of Minnesota and adjacent regions (No. 11). </em>Lubrecht &amp; Cramer Ltd.&nbsp; https://doi.org/10.2307/2422089</p>
<p>Schuster, R.M. (1966) <em>The Hepaticae and Anthocerotae of North America</em>, <em>East of the Hundredth Meridian I. </em>Columbia University Press, New York &amp; London.</p>
<p>Schuster, R.M. (1992) <em>The Hepaticae and Anthocerotae of North America, East of the Hundredth Meridian VI</em>. Field Museum of Natural History. Chicago.</p>
<p>Shaw, J. &amp; Renzaglia, K. (2004) Phylogeny and diversification of bryophytes. <em>American Journal of Botany </em>91 (10): 1557–1581.&nbsp; https://doi.org/10.3732/ajb.91.10.1557</p>
<p>Schwarz, U. &amp; Frahm, J.P. (2013) A contribution to the bryoflora of the Western Ghats in Karnataka State, India. <em>Polish Botanical Journal</em> 58 (2): 511–524.&nbsp; https://doi.org/10.2478/pbj-2013-0039</p>
<p>Singh, S.K. &amp; Singh, D.K. (2010) A catalogue of the liverworts and hornworts of Himachal Pradesh, India. <em>Archieve for Bryology </em>61.</p>
<p>Singh, D. &amp; Singh, D.K. (2017) Two new species of <em>Riccardia</em> (Aneuraceae, Marchantiophyta) from Eastern Himalaya, India with notes on the genus in Sikkim. <em>Taiwania</em> 62 (1): 33–42.</p>
<p>Singh, S.K., Bag, A.K. &amp; Bhattacharya, S.G. (2010) <em>Riccia </em>(Hepaticae: Ricciaceae) of West Bengal. <em>Taiwania </em>55 (2): 99–109.</p>
<p>Singh, S.K. &amp; Singh, D. (2018) A new species of the genus <em>Metzgeria</em> Raddi (Metzgeriaceae, Marchantiophyta) from India. <em>Cryptogamie, Bryologie</em> 39 (1): 47–53.&nbsp; https://doi.org/10.7872/cryb/v39.iss1.2018.47</p>
<p>Siregar, E.S., Ariyanti, N.S. &amp; Tjitrosoedirdjo, S.S. (2013) The Liverwort Genus <em>Marchantia </em>(Marchantiaceae) of Mount Sibayak North Sumatra, Indonesia. <em>Biotropia-The Southeast Asian Journal of Tropical Biology </em>20 (2): 73–80.&nbsp; https://doi.org/10.11598/btb.2013.20.2.327</p>
<p>So, M.L. (2002) <em>Metzgeria </em>(Hepaticae) in Australasia and the Pacific. <em>New Zealand journal of botany </em>40 (4): 603–627.&nbsp; https://doi.org/10.1080/0028825X.2002.9512818</p>
<p>So, M.L. (2003) The genus <em>Metzgeria</em> (Hepaticae) in Asia. <em>The Journal of the Hattori Botanical Laboratory</em> 94: 159–177.</p>
<p>So, M.L. (2004) <em>Metzgeria </em>(Metzgeriaceae, Marchantiophyta) in Africa. <em>New Zealand Journal of Botany </em>42 (2): 271–292.&nbsp; https://doi.org/10.1080/0028825X.2004.9512904</p>
<p>Söderström, L., Hagborg, A., von Konrat, M. &amp; Séneca, A. (2013) The Guaraní Land–checklist of hornworts (Anthocerotophyta) and liverworts (Marchantiophyta) of Paraguay. <em>Polish botanical journal </em>58 (1): 267–277.&nbsp; https://doi.org/10.2478/pbj-2013-0027</p>
<p>Söderström, L., Hagborg, A., von Konrat, M., Bartholomew-Began, S., Bell, D., Briscoe, L., Brown, E., Christine Cargill, D., Costa, D.P., Crandall-Stotler, B.J., Cooper, E.D., Dauphin, G., Engel, J.J., Feldberg, K., Glenny, D., Gradstein, S.R., He, X., Heinrichs, J., Hentschel, J., Ilkiu-Borges, A.L., Katagiri, T., Konstantinova, N.A., Larraín, J., Long, D.G., Nebel, M., Pócs, T., Puche, F., Reiner-Drehwald, E., Renner, M.A.M., Sass-Gyarmati, A., Schäfer-Verwimp, A., Segarra Moragues, J.G., Stotler, R.E., Sukkharak, P., Thiers, B.M., Uribe, J., Váňa, J., Villarreal, J.C., Wigginton, M., Zhang, L. &amp; Zhu, R.-L. (2016) World checklist of hornworts and liverworts. <em>PhytoKeys</em> 59: 1–828.&nbsp; https://doi.org/10.3897/phytokeys.59.6261</p>
<p>Soni, A., Kumar, A., Nath, V. &amp; Niveden, A. (2009) Genetic diversity of Indian liverwort <em>Plagiochasma appendiculatum</em> revealed by RAPD Marker. <em>Research Journal of Botany</em> 4 (3): 89–100.&nbsp; https://doi.org/10.3923/rjb.2009.89.100</p>
<p>Srivastava, S.C. &amp; Dixit, R. (1996) The genus <em>Cyathodium </em>Kunze. <em>Journal of the Hattori botanical laboratory </em>80: 149–215.</p>
<p>Srivastava, S.C. (1998) Distribution of hepaticae and anthocerotae in India. <em>In:</em> <em>Topics in bryology. </em>Allied Press: New Delhi. pp. 53–85.</p>
<p>Standley, P.C. (1915) Hepaticae of New Mexico. <em>The Bryologist </em>18 (6): 81–83.&nbsp; https://doi.org/10.1639/0007-2745(1915)18[81:HONM]2.0.CO;2</p>
<p>Stephani, F. (1898) <em>Species Hepaticarum </em>6. Genève &amp; Bale. pp. 784.</p>
<p>Stotler, R. &amp; Crandall-Stotler, B. (1977) A checklist of the liverworts and hornworts of North America. <em>Bryologist </em>80 (3): 405–428.&nbsp; https://doi.org/10.2307/3242017</p>
<p>Stotler, R., Crandall-Stotler, B.C. &amp; Cargill, C. (2016) Fossombroniaceae.<em> In: </em>Söderström, L., Hagborg, A. &amp; Konrat, M. von (Eds.) World checklist of hornworts and liverworts: 463467. <em>Phytokeys</em> 59.&nbsp; https://doi.org/10.3897/phytokeys.59.6261</p>
<p>Taylor, J., Hollingsworth, P.J. &amp; Bigelow, W.C. (1974) Scanning electron microscopy of liverwort spores and elaters. <em>Bryologist </em>77 (3)<em>:</em> 281–327. https://doi.org/10.2307/3241612</p>
<p>Udar, R. &amp; Chandra, V. (1964) <em>Exormotheca ceylonensis</em>-New record from India. <em>Current Science </em>33: 436–438.</p>
<p>Udar, R. &amp; Srivastava, S.C. (1968) Sporeling development in the genus Exormotheca. 1. <em>Exormotheca ceylonensis</em>. <em>Canadian Journal of Botany </em>46 (8): 1009–1012.&nbsp; https://doi.org/10.1139/b68-134</p>
<p>Underwood, L.M. (1895) Notes on our Hepaticae. III. The distribution of the North American Marchantiaceae. <em>Botanical Gazette </em>20 (2): 59–71.&nbsp; https://doi.org/10.1086/327151</p>
<p>Veltman, A., Potgieter, M.J. &amp; Van Wyk, A.E. (2000) <em>Metzgeria nicomariei</em>, a new species of Metzgeriaceae from South Africa. <em>South African Journal of Botany </em>66 (2): 112–114. https://doi.org/10.1016/S0254-6299(15)31071-1</p>
<p>Vithanage, P.W. (1970) A study of geomorphology and morphotechtonics of Ceylone.Proceedings of the 2nd seminar on geochemical prospecting methods and techniques. <em>UN Mineral Resources Development Series </em>38: 391–405.</p>
<p>Wachowiak, W., Bączkiewicz, A., Chudzińska, E. &amp; Buczkowska, K. (2007) Cryptic speciation in liverworts–a case study in the <em>Aneura pinguis </em>complex. <em>Botanical Journal of the Linnean Society </em>155 (2): 273–282.&nbsp; https://doi.org/10.1111/j.1095-8339.2007.00692.x</p>
<p>Wellman, C.H., Osterloff, P.L. &amp; Mohiuddin, U. (2003) Fragments of the earliest land plants. <em>Nature </em>425 (6955): 282–285.&nbsp; https://doi.org/10.1038/nature01884</p>
<p>Wheeler, J.A. (2000) Molecular phylogenetic reconstructions of the Marchantioid liverwort radiation. <em>The Bryologist </em>103 (2): 314–333.&nbsp; https://doi.org/10.1639/0007-2745(2000)103[0314:MPROTM]2.0.CO;2</p>
<p>Wigginton, M.J. (2001) A small collection of bryophytes from Ethiopia. <em>Tropical Bryology </em>20: 27–29.&nbsp; https://doi.org/10.11646/bde.20.1.4</p>
<p>Wikström, N., He-Nygrén, X. &amp; Shaw, A.J. (2009) Liverworts (Marchantiophyta). <em>The timetree of life</em>: 146–152.</p>
<p>Xiang, Y.L., Shu, L. &amp; Zhu, R.L. (2016) <em>Marchantia longii</em> (Marchantiaceae), a new species from northwestern Yunnan, China. <em>The Bryologist</em> 119 (3): 280–289.&nbsp; https://doi.org/10.1639/0007-2745-119.3.280</p>
<p>Zhang, Z.H., Peng, T., Li, X.N. &amp; Zhao, C.H. (2004) A study on the bryophytes of karst cave threshold at Kunming Area in Yunnan Province, PR China. <em>Carsologica Sinica </em>3: 013.</p>
<p>Zhaohui, Z. &amp; Pentecost, A. (2000) New and noteworthy bryophytes from the travertines of Guizhou and Sichuan, S.W. China. <em>Journal of Bryology </em>22 (1): 66–68.&nbsp; https://doi.org/10.1179/jbr.2000.22.1.66</p>
<p>Zubair, L. (2002) El Nino–southern oscillation influences on rice production in Sri Lanka. <em>International Journal of Climatology </em>22 (2): 249–260.&nbsp; https://doi.org/10.1002/joc.714</p>