Abstract
Themistoclesia diminuta, a new species of Ericaceae tribe Vaccinieae from western Colombia, is here described, discussed, and illustrated. It stands out among other Themistoclesia because of its subsessile flowers, pedicel articulate with calyx and obscure short pale green corollas with purple lobes.
References
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<p>Hooker, W.J. (1840) <em>Icones plantarum; or figures, with brief descriptive characters and remarks of new or rare plants, selected from the author’s herbarium Vol. 3.</em> Longman, Rees, Orme, Brown, Green, & Longman. London, 99 pp.</p>
<p>Klotzsch, J.F. (1851) Studen über die natürliche klasse Bicornes Linné. <em>In</em>: Schlechtendal, D.F.L. (Ed.) <em>Linnaea</em> 24 (1): 1–88.</p>
<p>Kron, K.A., Powell, E.A. & Luteyn, J.L. (2002) Phylogenetic relationships within the blueberry tribe (Vaccinieae, Ericaceae) based on sequence data from <em>matK </em>and nuclear ribosomal ITS regions, with comments on the placement of <em>Satyria</em>. <em>American Journal of Botany</em> 89: 327–336. https://doi.org/10.3732/ajb.89.2.327</p>
<p>Luteyn, J.L. (2002) Diversity, adaptation, and endemism in neotropical Ericaceae: biogeographical patterns in the Vaccinieae. <em>In</em>: Young, K., Ulloa Ulloa, C., Luteyn, J.L. & Knapp, S. (Eds.) Plant evolution and endemism in Andean South America. <em>Botanical Review</em> 68 (1): 55–87. https://doi.org/10.1663/0006-8101(2002)068[0055:DAAEIN]2.0.CO;2</p>
<p>Luteyn, J.L. & Pedraza-Peñalosa, P. (2012) <em>Blueberry relatives of the New World tropics (Ericaceae)</em>. The New York Botanical Garden, Bronx, New York. Available from: http://sweetgum.nybg.org/ericaceae/index.php (accessed 8 February 2022).</p>
<p>Luteyn, J.L. & Pedraza-Peñalosa, P. (2013) Nomenclature, taxonomy, and conservation of the Neotropical genus <em>Sphyrospermum</em> (Ericaceae: Vaccinieae), including five new species from Colombia, Ecuador and Peru. <em>Phytotaxa</em> 79: 1–29. https://doi.org/10.11646/phytotaxa.79.1.1</p>
<p>Pedraza-Peñalosa, P. (2008) Three new species of <em>Disterigma</em> (Ericaceae: Vaccinieae) from western Colombia, with comments on morphological terminology. <em>Brittonia</em> 60 (1): 1–10. https://doi.org/10.1007/s12228-008-9006-3</p>
<p>Pedraza-Peñalosa, P. (2015a) New blueberry and <em>mortiño </em>relatives (Ericaceae) from northwestern Colombia. <em>PhytoKeys</em> 49: 33–58. https://doi.org/10.3897/phytokeys.49.8383</p>
<p>Pedraza-Peñalosa, P. (2015b) Three new species of <em>Psammisia</em> s.l. (Ericaceae: Vaccinieae), blueberry relatives endemic to Colombia. <em>Brittonia</em> 68 (1): 25–36. https://doi.org/10.1007/s12228-015-9388-y</p>
<p>Pedraza-Peñalosa, P. (2016) Ericaceae. <em>In</em>: Bernal, R., Gradstein, S.R. & Celis, M. (Eds.) <em>Catálogo de plantas y líquenes de Colombia. Vol. I.</em> Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá, pp. 1188–1212.</p>
<p>Pedraza-Peñalosa, P. & Luteyn, J.L. (2010) New species of Ericaceae (Vaccinieae) from the Andes of Bolivia and Peru. <em>Brittonia</em> 62 (1): 39–56. https://dx.doi.org/10.1007/s12228-009-9096-6</p>
<p>Pedraza-Peñalosa, P., Salinas, N.R., Virnig, A.L.S. & Wheeler, W.C. (2015) Preliminary phylogenetic analysis of the Andean clade and the placement of new Colombian blueberries (Ericaceae, Vaccinieae). <em>PhytoKeys</em> 49: 13–31. https://doi.org/10.3897/phytokeys.49.8622</p>
<p>Rangel-Ch., O. (2000) La región paramuna y franja aledaña en Colombia. <em>In</em>: Rangel-Ch., J.O. (Ed.) <em>La región de vida paramuna. Colombia diversidad biótica III. </em>Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá, pp. 1–23.</p>
<p>Saint-Hilaire, J.H.J. (1805) Ericaceae.<em> In:</em> <em>Exposition des Familles Naturelles et de la germination des plantes</em> 1. pp. 1–512.</p>
<p>Sleumer, H. (1934) Ericaceae americanae novae vel minus cognitae. <em>Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem</em> 12 (112): 119–140. https://doi.org/10.2307/3995031</p>
<p>Sleumer, H. (1935) Ericaceae americanae novae vel minus cognitae II. <em>Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem</em> 12 (113): 277–294. https://doi.org/10.2307/3994889</p>
<p>Sleumer, H. (1941) Vaccinioideen-Studien. <em>Botanischen Jahrbücher fur Systematik, Pflanzengeschichte und Pflanzengeographie</em> 71 (4): 375–510.</p>
<p>Smith, A.C. (1936) Studies of South American Plants-V Additional Notes on Thibaudieae. <em>Bulletin of the Torrey Botanical Club</em> 63 (6): 307–316. https://doi.org/10.2307/2481157</p>
<p>Smith, A.C. (1950) Studies of South American Plants XII. <em>Contributions from the United States National Herbarium</em> 29 (8): 317–393.</p>
<p>Standley, P.C. (1938) Flora of Costa Rica. <em>Publications of the Field Museum of Natural History, Botanical Series</em> 18 (3): 783–1133.</p>
<p>Cronquist, A. (1978) Once again, what is a species? <em>In:</em> Romberger, J.A. (Ed.) <em>Biosystematics in agriculture.</em> Al-lanheld Osmun, Montclair, pp. 3–20.</p>
<p>Hooker, W.J. (1840) <em>Icones plantarum; or figures, with brief descriptive characters and remarks of new or rare plants, selected from the author’s herbarium Vol. 3.</em> Longman, Rees, Orme, Brown, Green, & Longman. London, 99 pp.</p>
<p>Klotzsch, J.F. (1851) Studen über die natürliche klasse Bicornes Linné. <em>In</em>: Schlechtendal, D.F.L. (Ed.) <em>Linnaea</em> 24 (1): 1–88.</p>
<p>Kron, K.A., Powell, E.A. & Luteyn, J.L. (2002) Phylogenetic relationships within the blueberry tribe (Vaccinieae, Ericaceae) based on sequence data from <em>matK </em>and nuclear ribosomal ITS regions, with comments on the placement of <em>Satyria</em>. <em>American Journal of Botany</em> 89: 327–336. https://doi.org/10.3732/ajb.89.2.327</p>
<p>Luteyn, J.L. (2002) Diversity, adaptation, and endemism in neotropical Ericaceae: biogeographical patterns in the Vaccinieae. <em>In</em>: Young, K., Ulloa Ulloa, C., Luteyn, J.L. & Knapp, S. (Eds.) Plant evolution and endemism in Andean South America. <em>Botanical Review</em> 68 (1): 55–87. https://doi.org/10.1663/0006-8101(2002)068[0055:DAAEIN]2.0.CO;2</p>
<p>Luteyn, J.L. & Pedraza-Peñalosa, P. (2012) <em>Blueberry relatives of the New World tropics (Ericaceae)</em>. The New York Botanical Garden, Bronx, New York. Available from: http://sweetgum.nybg.org/ericaceae/index.php (accessed 8 February 2022).</p>
<p>Luteyn, J.L. & Pedraza-Peñalosa, P. (2013) Nomenclature, taxonomy, and conservation of the Neotropical genus <em>Sphyrospermum</em> (Ericaceae: Vaccinieae), including five new species from Colombia, Ecuador and Peru. <em>Phytotaxa</em> 79: 1–29. https://doi.org/10.11646/phytotaxa.79.1.1</p>
<p>Pedraza-Peñalosa, P. (2008) Three new species of <em>Disterigma</em> (Ericaceae: Vaccinieae) from western Colombia, with comments on morphological terminology. <em>Brittonia</em> 60 (1): 1–10. https://doi.org/10.1007/s12228-008-9006-3</p>
<p>Pedraza-Peñalosa, P. (2015a) New blueberry and <em>mortiño </em>relatives (Ericaceae) from northwestern Colombia. <em>PhytoKeys</em> 49: 33–58. https://doi.org/10.3897/phytokeys.49.8383</p>
<p>Pedraza-Peñalosa, P. (2015b) Three new species of <em>Psammisia</em> s.l. (Ericaceae: Vaccinieae), blueberry relatives endemic to Colombia. <em>Brittonia</em> 68 (1): 25–36. https://doi.org/10.1007/s12228-015-9388-y</p>
<p>Pedraza-Peñalosa, P. (2016) Ericaceae. <em>In</em>: Bernal, R., Gradstein, S.R. & Celis, M. (Eds.) <em>Catálogo de plantas y líquenes de Colombia. Vol. I.</em> Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá, pp. 1188–1212.</p>
<p>Pedraza-Peñalosa, P. & Luteyn, J.L. (2010) New species of Ericaceae (Vaccinieae) from the Andes of Bolivia and Peru. <em>Brittonia</em> 62 (1): 39–56. https://dx.doi.org/10.1007/s12228-009-9096-6</p>
<p>Pedraza-Peñalosa, P., Salinas, N.R., Virnig, A.L.S. & Wheeler, W.C. (2015) Preliminary phylogenetic analysis of the Andean clade and the placement of new Colombian blueberries (Ericaceae, Vaccinieae). <em>PhytoKeys</em> 49: 13–31. https://doi.org/10.3897/phytokeys.49.8622</p>
<p>Rangel-Ch., O. (2000) La región paramuna y franja aledaña en Colombia. <em>In</em>: Rangel-Ch., J.O. (Ed.) <em>La región de vida paramuna. Colombia diversidad biótica III. </em>Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá, pp. 1–23.</p>
<p>Saint-Hilaire, J.H.J. (1805) Ericaceae.<em> In:</em> <em>Exposition des Familles Naturelles et de la germination des plantes</em> 1. pp. 1–512.</p>
<p>Sleumer, H. (1934) Ericaceae americanae novae vel minus cognitae. <em>Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem</em> 12 (112): 119–140. https://doi.org/10.2307/3995031</p>
<p>Sleumer, H. (1935) Ericaceae americanae novae vel minus cognitae II. <em>Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem</em> 12 (113): 277–294. https://doi.org/10.2307/3994889</p>
<p>Sleumer, H. (1941) Vaccinioideen-Studien. <em>Botanischen Jahrbücher fur Systematik, Pflanzengeschichte und Pflanzengeographie</em> 71 (4): 375–510.</p>
<p>Smith, A.C. (1936) Studies of South American Plants-V Additional Notes on Thibaudieae. <em>Bulletin of the Torrey Botanical Club</em> 63 (6): 307–316. https://doi.org/10.2307/2481157</p>
<p>Smith, A.C. (1950) Studies of South American Plants XII. <em>Contributions from the United States National Herbarium</em> 29 (8): 317–393.</p>
<p>Standley, P.C. (1938) Flora of Costa Rica. <em>Publications of the Field Museum of Natural History, Botanical Series</em> 18 (3): 783–1133.</p>