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Published: 2023-01-13

Typifications of Andean Arenaria names (Caryophyllaceae), with a nomen novum

Botanic Garden and Botanical Museum Berlin (BGBM), Freie Universität Berlin, Königin-Luise-Str. 6–8, 14195 Berlin, Germany. Instituto Científico Michael Owen Dillon, Av. Jorge Chávez 610, Cercado, Arequipa, Peru. Instituto de Ciencia y Gestión Ambiental de la Universidad Nacional de San Agustín de Arequipa. Calle San Agustín 108, Arequipa-Peru.
Ce.R.S.I.Te.S., Viale XXIV Maggio 7, 04100 Latina, Italy
Alsine Arenaria new combination new name Scleranthus typification Eudicots


The nomenclature of the names Alsine weberbaueri, Arenaria musciformis, and Scleranthus peruvianus is investigated. The names were typified on specimens preserved at HOXA, MOL, and P. All these names are to be included in Arenaria. Concerning Alsine weberbaueri, a new combination (Arenaria weberbaueri) is proposed, whereas for Arenaria humboldtiana, a new name is provided for Scleranthus peruvianus since a previous Arenaria peruviana by Poeppig ex Rohrbach was validly published (Art. 53.1 of ICN).


  1. Bernal, R., Gradstein, S.R. & Celis, M. (eds.) (2019) Catálogo de plantas y líquenes de Colombia. Available from: (accessed 29 November 2022).

  2. Bittrich, V. (1993) Caryophyllaceae. In: Kubitzki, K, Rohwer, J.G. (eds.) The families and genera of vascular plants, Vol. 2. Springer, Berlin, pp. 206–236.

  3. Brako, L. & Zarucchi, J. (1993) Catalogue of the Flowering Plants and Gymnosperms of Peru. Monographs in Systematic Botany from the Missouri Botanical Garden 45: 1–1286.

  4. Briceño, B. & Morillo, G. (2002) Catálogo abreviado de las plantas con flores de los páramos de Venezuela. Parte I. Dicotiledóneas (Magnoliopsida). Acta Botánica Venezuelica 25: 1–42.

  5. Chen, S.L. & Rabeler, R.K. (2001) Stellaria L. In: Wu, Z.Y., Raven, P.H. & Hong, D.Y. (eds.) Flora of China, vol. 26. Science Press, & St Louis, Beijing, pp. 11–29.

  6. Conti, F., Bartolucci, F., Iamonico, D. & Del Guacchio, E. (2014) A new combination in Mcneillia (Caryophyllaceae) for the Italian flora. Phytotaxa 170 (2): 139–140.

  7. Dillenberger, M.S. & Kadereit, J.W. (2014) Maximum polyphyly: Multiple origins and delimitation with plesiomorphic characters require a new circumscription of Minuartia (Caryophyllaceae). Taxon 63 (1): 64–88.

  8. Greenberg, A.K. & Donoghue, M.J. (2011) Molecular systematics and character evolution in Caryophyllaceae. Taxon 60 (6): 1637–1652.

  9. Hernández-Ledesma, P., Berendsohn, W.G., Borsch, T., von Mering, S., Akhani, H., Arias, S., Castañeda-Noa, I., Eggli, U., Eriksson, R., Flores-Olvera, H., Fuentes-Bazán, S., Kadereit, G., Klak, C., Korotkova, N., Nyffeler, R., Ocampo, G., Ochoterena, H., Oxelman, B., Rabeler, R.K., Sanchez, A., Schlumpberger, B.O. & Uotila, P. (2015) A taxonomic backbone for the global synthesis of species diversity in the angiosperm order Caryophyllales. Willdenowia 45: 281–383.

  10. Hiepko, P. (1987) The collections of the Botanical Museum Berlin Dahlem (B) and their history. In: Scholtz, H. (Ed.) Botany in Berlin. Englera 7: 219–252.

  11. Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008) Nuevo Catálogo de la Flora Vascular de Venezuela. Fundación Instituto Botánico de Venezuela, Caracas, 859 pp.

  12. HUH Index of botanists (2013 onwards) Index of botanists, Harward University Herbaria & Libraries. Available from: (accessed 29 November 2022).

  13. Hupp, N., Llambí, L.D., Ramírez, L. & Callaway, R.M. (2017) Alpine cushion plants have species–specific effects on microhabitat and community structure in the tropical Andes. Journal of Vegetation Science 28 (5): 928–938.

  14. Iamonico, D. (2013) Taxonomical and chorological study on the Central Mediterranean Basin endemic Arenaria bertolonii Fiori & Paol. (Caryophyllaceae). Plant Biosystems 147 (4): 923–930.

  15. Iamonico, D. (2014) Arenaria tenuifolia versus Arenaria hybrida (Caryophyllaceae): nomenclatural study and taxonomic implications. Phytotaxa 173 (3): 235–240.

  16. Iamonico, D. (2016) Nomenclatural notes on four Linnaean names in Arenaria (Caryophyllaceae). Taxon 65 (3): 610–616.

  17. Iamonico, D. (2019) Taxonomical, chorological and ecological notes on the Eastern Alps endemic Arenaria huteri (Caryophyllaceae-Alsinoideae). Phyton (Horn, Austria) 59 (1): 11–19.

  18. Iamonico, D. (2022) Notas nomenclaturales sobre el género Arenaria (Caryophyllaceae) en Chile. Darwiniana nueva serie 10 (1): 187–192.

  19. Linnaeus, C. (1753) Species plantarum, vol. 1. Laurentii Salvii, Holmiae, 560 pp.

  20. Llambí, L.D., Hupp, N., Saez, A. & Callaway, R. (2018) Reciprocal interactions between a facilitator, natives, and exotics in tropical alpine plant communities. Perspectives in Plant Ecology, Evolution and Systematics 30: 82–88.

  21. MacBride, J.F. (1936) Caryophyllaceae Juss. Flora of Peru. Publications of the Field Museum of Natural History. Botanical series 13 (2): 578–639.

  22. Molinari, E. (2016) Further nomenclatural changes regarding Peruvian endemics. Polish Botanical Journal 61: 275–277. https://10.1515/pbj-2016-0021

  23. Muschler, R. (1911) Caryophyllaceae andinae. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 45: 441–461.

  24. POWO (2022a) Plants of the World Online. Alsine L. Available from: (accessed 29 November 2022).

  25. POWO (2022b) Plants of the World Online. Alsine L. Available from: (accessed 29 November 2022).

  26. POWO (2022c) Plants of the World Online. Scleranthus peruvianus Muschl. Available from: (accessed 29 November 2022).

  27. POWO (2022d) Plants of the World Online. Scleranthus annuus L. Available from: (accessed 29 November 2022).

  28. Rabeler, R.K. & Hartmann, R.L. (2005) Caryophyllaceae Juss. In: Flora of North America Editorial Committee (eds.) Flora of North America North of Mexico, vol. 5. Oxford University Press, New York & Oxford, U.S.A., pp. 3–8.

  29. Rohrbach, P. (1871–1873[1872]) Beiträge zur Systematik der Caryophillinen. Linnaea: Ein Journal für die Botanik in ihrem ganzen Umfange 37: 183–312.

  30. Sadeghian, S., Zarre, S., Rabeler, R.K. & Heubl, G. (2015) Molecular phylogenetic analysis of Arenaria (Caryophyllaceae: tribe Arenarieae) and its allies inferred from nuclear DNA internal transcribed spacer and plastid DNA rps16 sequences. Botanical Journal of the Linnean Society 178 (4): 648–669.

  31. Sleumer, H. (1949) The Botanical Gardens and Museum at Berlin-Dahlem. Kew Bulletin 4 (2): 172–175.

  32. Stafleu, A.F. & Cowan, R.S. (1983) Taxonomic literature, 2nd ed., vol. 4 (P–Sak). Bohn, Scheltema & Holtema, Utrecht, 1214 pp.

  33. Stafleu, A.F. & Cowan, R.S. (1986) Taxonomic literature, 2nd ed., vol. 6 (Sti–Vuy). Bohn, Scheltema & Holtema, Utrecht, 926 pp.

  34. Stafleu, A.F. & Cowan, R.S. (1988) Taxonomic literature, 2nd ed., vol. 7 (W–Z). Bohn, Scheltema & Holtema, Utrecht, 653 pp.

  35. Thiers, B. (2022) [continuously updated] Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. (accessed 29 November 2022).

  36. Triana, J.J. & Planchon, J.E. (1862) Prodromus Florae Novo-Granatensis ou énumération des plantes de la Nuovelle-Grenade vec description des espècies nouvelles. Annales de Sciences Naturelles, Botanique, série 4 17: 1–382.

  37. Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress, Shenzhen, China, July 2017. Regnum Vegetabile 159: 1–254.

  38. Ulloa Ulloa, C., Acevedo-Rodríguez, P., Beck, S.G., Belgrano, M.J., Bernal, R., Berry, P.E., Brako, L., Celis, M., Davidse, G., Gradstein, S.R., Hokche, O., León, B., León-Yánez, S., Magill, R.E., Neill, D.A., Nee, M.H., Raven, P.H., Stimmel, H., Strong, M.T., Villaseñor Ríos, J.L., Zarucchi, J.L., Zuloaga, F.O. & Jørgensen, P.M. (2017) An integrated assessment of vascular plants species of the Americas. Science 358: 1614–1617.

  39. Walters, S.M. (Ed.) (1993) Caryophyllaceae Juss. In: Tutin, T.G., Burges, N.A., Chater, A.O, Edmondson, J.R., Heywood, V.H., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A. (eds.) Flora Europaea, ed. 2, vol. 1. Cambridge University Press, Cambridge, pp. 139.

  40. West, J.G. & Garnock-Jones, P.J. (1986) Evolution and biogeography of Scleranthus (Caryophyllaceae) in Australasia. In: Barlow, B.A (Eds.) Flora and Fauna of Alpine Australasia. Ages and Origins. Sciro in association with Australian Systematic Botany Society. Australia. pp. 435–450.