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Type: Editorial
Published: 2020-02-17
Page range: 8–13
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In Memoriam: Dr. Rut Collado de la Peña (1967–2015)

General Secretary, ISAO Illinois Natural History Survey, Champaign, Illinois 61820, USA


Our dear friend and colleague, Dr. Rut Collado de la Peña, passed away on 14 November 2015, in A Coruña, Galicia, Spain, at the age of 48.
She was an associate professor ("Profesora Contratada Doctora") and member of the Science Faculty within the Department of Animal Biology, Plant Biology and Ecology at the Universidade da Coruña, a Spanish public university in the city of A Coruña, in Galicia, northwestern Spain.


  1. Collado, R. & Martínez-Ansemil, E. (1991) Les Oligochètes aquatiques de la Péninsule Ibérique: premières données sur le Portugal. Bulletin de la Société d'Histoire Naturelle,Toulouse, 127, 57–69.

    Collado, R., Martínez-Ansemil, E. & Giani, N. (1993) Les Oligochètes aquatiques de la Péninsule Ibérique: description de Stylodrilus curvithecus n.sp. (Lumbriculidae) et de Mesenchytraeus lusitanicus n. sp. et redescription de Cognettia hibernica Healy (Enchytraeidae). Annales de Limnologie, Toulouse, 29, 129–138.

    Martínez-Ansemil, E. & Collado, R. (1996) Distribution patterns of aquatic oligochaetes inhabiting watercourses in the Northwestern Iberian Peninsula. Hydrobiologia, 334, 73–83.

    Martínez-Ansemil, E. & Collado, R. (1996) Two new species of freshwater Oligochaeta from the North-west Iberian Peninsula: Krenedrilus realis sp. nov. (Tubificidae) and Cernosvitoviella bulboducta sp. nov. (Enchytraeidae). Journal of Zoology, London, 240, 363–370.

    Collado, R. & Martínez-Ansemil, E. (1996) Inventario dos Oligoquetos acuáticos de Galicia (Annelida: Oligochaeta). Cadernos da Área de Ciencias Biolóxicas (Inventarios). Seminario de Estudos Galegos, XIV. Edicións do Castro, O Castro-Sada, A Coruña, 38 pp. [in Galician, with French and English summary]

    Collado, R., Kasprzak, P. & Schmelz, R. (1999) Oligochaeta and Aphanoneura in two Northern German hardwater lakes of different trophic state. Hydrobiologia, 406, 143–148.

    Schmelz, R.M. & Collado, R. (1999) Enchytraeus luxuriosus sp. nov., a new terrestrial oligochaete species (Enchytraeidae, Clitellata, Annelida). carolinea, 57, 93–100.

    Schmelz, R.M., Collado, R. & Dózsa-Farkas, K. (1999) New record of Fridericia pyrenaica Giani, 1979, with complements to the original description. Bulletin de la Société d'Histoire Naturelle, Toulouse, 135, 71–75.

    Schmelz, R.M., Collado, R. & Oestreich, M. (1999) The Enchytraeus buchholzi/christenseni species complex - outline of a problem and first steps towards a solution. In: Schmelz, R.M. & Sühlo, K. (Eds.), Newsletter on Enchytraeidae No. 6. Proceedings of the 3rd International Symposium on Enchytraeidae, Osnabrück, Germany. Universitätsverlag Rasch, Osnabrück, Germany, pp. 31–39.

    Collado, R., Schmelz, R.M., Moser, T. & Römbke, J. (1999) Enchytraeid reproduction test (ERT): Sublethal responses of two Enchytraeus species (Oligochaeta) to toxic chemicals. Pedobiologia, 43, 625–629.

    Collado, R. & Schmelz, R.M. (2000) Pedonais crassifaucis n.gen., n.sp. (Naididae) and Bothrioneurum righii (Tubificidae), two new soil-dwelling species of "aquatic" oligochaetes (Clitellata, Annelida) from Central Amazonia. Amazoniana, 16, 223–235.

    Collado, R. & Schmelz, R.M. (2000) Pristina silvicola and Pristina terrena spp. nov., two new soil-dwelling species of Naididae (Oligochaeta, Annelida) from the tropical rain forest near Manaus, Brazil, with comments on the genus Pristinella. The Journal of Zoology, 251, 509–516.

    Schmelz, R.M., Collado, R. & Myohara, M. (2000) A taxonomical study of Enchytraeus japonensis (Enchytraeidae, Oligochaeta): Morphological and biochemical comparisons with E. bigeminus. Zoological Science, 17, 505–516.[505:ATSOEJ]2.0.CO;2

    Collado, R. & Schmelz, R.M. (2001) Descriptions of three Pristina species (Naididae, Clitellata) from Amazonian forest soils, including P. marcusi sp. nov. Hydrobiologia, 463, 1–11.

    Collado, R. & Schmelz, R.M. (2001) Oligochaete distribution patterns in two German hardwater lakes of different trophic state. Limnologica, Berlin, 31, 317–328.

    Collado, R. & Schmelz, R.M. (2002) Pristina trifida sp. nov., a new soil-dwelling microannelid (Oligochaeta: Naididae) from Amazonian forest soils, with comments on species recognition in the genus. Zootaxa, 118, 1–14.

    Römbke, J., Collado, R. & Schmelz, R.M. (2005) Oligochaetes (Clitellata) of the Mata Atlântica (Parana, Brazil): first results of the SOLOBIOMA project. Proceedings of the Estonian Academy of Sciences: Biology, Ecology, 54, 302–309.

    Römbke, J., Collado, R. & Schmelz, R.M. (2007) Abundance, distribution, and indicator potential of enchytraeid genera (Enchytraeidae, Clitellata) in secondary forests and pastures of the Mata Atlântica (Paraná, Brazil). Acta Hydrobiologica Sinica, 31 Suppl., 139–150.

    Schmelz, R.M., Collado de la Peña, R.M. & Römbke, J. (2009) Benefits from ecological study methods to taxonomy of enchytraeids in southern Mata Atlântica. Pesquisa agropecuaria brasileira, Brasilia, 44, 861–867.

    Schmelz, R.M. & Collado, R. (2005) Achaeta becki sp. nov. (Oligochaeta: Enchytraeidae) from Amazonian forest soils. Zootaxa, 1084, 49–57.

    Schmelz, R.M. & Collado, R. (2005) Fridericia larix sp. nov. (Enchytraeidae, Oligochaeta) from Irish soils. Organisms, Diversity & Evolution, 5, 85–88.

    Schmelz, R.M. & Collado, R. (2007) Revision of Hemienchytraeus stephensoni (Cognetti, 1927) (Enchytraeidae, Oligochaeta, Annelida). Folia Facultatis scientiarum naturalium Universitatis Masarykianae Brunensis, Biologiai, 110, 67–85.

    Schmelz, R.M., Collado, R. & Briones, M.J.I. (2008) Notes on Marionina filiformis (Enchytraeidae, Oligochaeta, Annelida). Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg (Neue Folge), 44, 23–35.

    Schmelz, R.M., Collado, R. & Römbke, J. (2008) Mata Atlântica enchytraeids (Paraná, Brazil): The genus Achaeta (Oligochaeta, Enchytraeidae). Zootaxa, 1809, 1–35.

    Schmelz, R.M. & Collado, R.M. (2008) A type-based redescription of Pachydrilus georgianus Michaelsen, 1888, the type species of Marionina Michaelsen, 1890, with comments on Christensenidrilus Dózsa-Farkas & Convey, 1997 (Enchytraeidae, "Oligochaeta", Annelida). Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg (Neue Folge), 44, 7–22.

    Collado, R. & de Mendoza, G. (2009) Environmental factors and distribution of littoral oligochaetes in Pyrenean lakes. Advances in Limnology, 62, 215–244.

    Parapar, J., Martínez-Ansemil, E., Caramelo, C., Collado, R. & Schmelz, R. (2009) Polychaetes and oligochaetes associated with intertidal rocky shores in a semi-enclosed industrial and urban embayment, with the description of two new species. Helgoland Marine Research, 63, 293–308.

    Schmelz, R.M. & Collado, R. (2010) A guide to European terrestrial and freshwater species of Enchytraeidae (Oligochaeta). Soil Organisms, 82, 1–176.

    Schmelz, R.M., Collado, R. & Römbke, J. (2011) Mata Atlântica enchytraeids (Enchytraeidae, Oligochaeta): Description of a new genus, Xetadrilus gen. nov., with three new species, and four new species of Guaranidrilus Černosvitov. Zootaxa, 2838, 1–29.

    Schmelz, R.M. & Collado, R. (2012) Guide to European terrestrial and freshwater species of Enchytraeidae (Oligochaeta): first supplement. vTI Agriculture and Forestry Research, Special Issue, 357, 53–66.

    Collado, R., Haß-Cordes, E. & Schmelz, R.M. (2012) Microtaxonomy of fragmenting Enchytraeus species using molecular markers, with a comment on species complexes in enchytraeids. Turkish Journal of Zoology, 36, 85–94.

    Schmelz, R.M. & Collado, R. (2012) An updated checklist of currently accepted species of Enchytraeidae (Oligochaeta). vTI Agriculture and Forestry Research, Special Issue, 357, 67–87.

    Schmelz, R.M. & Collado, R. (2013) 'Cejkaian tubules' in the posterior midgut of terrestrial Enchytraeidae (Oligochaeta). Soil Organisms, 85, 113–122.

    Schmelz, R.M. & Collado, R. (2013) Enchytraeidae (Oligochaeta, Annelida) from a field site in Portugal, with the description of five new species and a redescription of Enchylea heteroducta Nielsen & Christensen, 1963. Zootaxa, 3647 (2), 307–328.

    Schmelz, R.M., Niva, C.C., Römbke, J. & Collado, R. (2013) Diversity of terrestrial Enchytraeidae (Oligochaeta) in Latin America: Current knowledge and future research potential. Applied Soil Ecology, 69, 13–20.

    Schmelz, R.M. & Collado, R. (2015) Checklist of taxa of Enchytraeidae (Oligochaeta): an update. Soil Organisms, 87, 149–152.

    Schmelz, R.M., Jocque, M. & Collado, R. (2015) Microdrile Oligochaeta in bromeliad pools of a Honduran cloud forest. Zootaxa, 3947 (4), 508–526.

    Römbke, J., Collado, R., Höfer, H., Ottermanns, R., Raub, F., Ross-Nickoll, M. & Schmelz, R.M. (2015) Species diversity of Enchytraeidae (Oligochaeta) in pastures, regenerating secondary forests, and old-growth forests in the southern Mata Atlântica. Soil Organisms, 87, 101–120.