Kohlmann, B. and Solís, Ángel 2012.

New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama

. Zootaxa. 3193, 1 (Feb. 2012), 28–52. DOI:https://doi.org/10.11646/zootaxa.3193.1.2.