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Johnson, C.B., S. H., E.Q., Anuar, S., Muin, M.A., Wood, Jr., P.L., Grismer, J.L., Greer, L.F., Onn, C.K., Ahmad, N., Bauer, A.M. and Grismer, L.L. 2012. Phylogeography, geographic variation, and taxonomy of the Bent-toed Gecko Cyrtodactylus quadrivirgatus Taylor, 1962 from Peninsular Malaysia with the description of a new swamp dwelling species. Zootaxa. 3406, 1 (Aug. 2012), 39–58. DOI:https://doi.org/10.11646/zootaxa.3406.1.3.