Cardoso, Y.P., Almirón, A., Casciotta, J., Aichino, D., Lizarralde, M.S. and Montoya-Burgos, J.I. 2012. Origin of species diversity in the catfish genus Hypostomus (Siluriformes: Loricariidae) inhabiting the Paraná river basin, with the description of a new species. Zootaxa. 3453, 1 (Sep. 2012), 69–83. DOI:https://doi.org/10.11646/zootaxa.3453.1.5.