Sulaiman, Z.H. and Mayden, R.L. 2012. Cypriniformes of Borneo (Actinopterygii, Otophysi): An Extraordinary Fauna for Integrated Studies on Diversity, Systematics, Evolution, Ecology, and Conservation. Zootaxa. 3586, 1 (Dec. 2012), 359–376. DOI:https://doi.org/10.11646/zootaxa.3586.1.34.