Fominykh, D.D., Zamotajlov, A.S. and Titarenko, A.Y. 2016.

Further contribution to the knowledge of Georgian species of the Carabus Linnaeus, 1758 subgenus Neoplectes Reitter, 1885, with description of a new subspecies of C. titarenkoi Zamotajlov & Fominykh, 2014 (Coleoptera: Carabidae: Carabini)

. Zootaxa. 4179, 3 (Oct. 2016), 561–599. DOI:https://doi.org/10.11646/zootaxa.4179.3.10.