González-Durán, G.A., Targino, M., Rada, M. and Grant, T. 2017.

Phylogenetic relationships and morphology of the Pristimantis leptolophus species group (Amphibia: Anura: Brachycephaloidea), with the recognition of a new species group in Pristimantis Jiménez de la Espada, 1870

. Zootaxa. 4243, 1 (Mar. 2017), 42–74. DOI:https://doi.org/10.11646/zootaxa.4243.1.2.