Protogino, L.C., Miquelarena, A.M. and López, H.L. 2006. A new species of Astyanax (Teleostei, Characiformes, Characidae), with breeding tubercles, from the Paraná and Uruguay river basins. Zootaxa. 1297, 1 (Aug. 2006), 1–16. DOI:https://doi.org/10.11646/zootaxa.1297.1.1.