Kohlmann, B. 2007.

Biodiversity, conservation, and hotspot atlas of Costa Rica: a dung beetle perspective (Coleoptera: Scarabaeidae: Scarabaeinae)

. Zootaxa. 1466, 1 (May 2007), 68. DOI:https://doi.org/10.11646/zootaxa.1466.1.7.