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Venegas, P.J., Barboza, A.C., Riva, I.D.L. and Padial, J.M. 2018.

A new species of Phrynopus from the northeastern Andes of Peru, its phylogenetic position, and notes on the relationships of Holoadeninae (Anura: Craugastoridae)

. Zootaxa. 4446, 4 (Jul. 2018), 501–524. DOI:https://doi.org/10.11646/zootaxa.4446.4.5.