Mendelson Iii, J.R., Mulcahy, D.G., Williams, T.S. and Sites Jr., J.W. 2011.

A phylogeny and evolutionary natural history of mesoamerican toads (Anura: Bufonidae: Incilius) based on morphology, life history, and molecular data

. Zootaxa. 3138, 1 (Dec. 2011), 1–34. DOI:https://doi.org/10.11646/zootaxa.3138.1.1.