Rabemananjara, F.C., Crottini, A., Chiari, Y., Andreone, F., Glaw, F., Duguet, R., Bora, P., Ramilijaona, O.R. and Vences, M. 2007.

Molecular systematics of Malagasy poison frogs in the Mantella betsileo and  M. laevigata species groups

. Zootaxa. 1501, 1 (Jun. 2007), 31–44. DOI:https://doi.org/10.11646/zootaxa.1501.1.2.