Coloma, L.A., Lötters, S., Duellman, W.E. and Miranda-Leiva, A. 2007.

A taxonomic revision of Atelopus pachydermus, and description of two new (extinct?) species of Atelopus from Ecuador (Anura: Bufonidae)

. Zootaxa. 1557, 1 (Aug. 2007), 1–32. DOI:https://doi.org/10.11646/zootaxa.1557.1.1.