Valencia, J.H., Valladares-Suntasig, F., Tipantiza-Tuguminago, L. and Dueñas, M.R. 2019. A new species of terrestrial-breeding frog of the genus Pristimantis (Anura: Terrarana: Craugastoridae) from the eastern Andean slopes of the southern Ecuador. Zootaxa. 4658, 3 (Aug. 2019), 509–525. DOI:https://doi.org/10.11646/zootaxa.4658.3.4.