Maddison, D.R., Toledano, L., Sallenave, S. and Roig-Juñent, S. 2013. Phylogenetic relationships of the South American ground beetle subgenus Chilioperyphus Jeannel (Coleoptera: Carabidae: Trechinae: Bembidiini: Bembidion Latreille). Zootaxa. 3636, 4 (Apr. 2013), 547–560. DOI:https://doi.org/10.11646/zootaxa.3636.4.3.