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Angulo, A. and Gracian-Negrete, J.M. 2013. A new species of Brycon (Characiformes: Characidae) from Nicaragua and Costa Rica, with a key to the lower Mesoamerican species of the genus. Zootaxa. 3731, 2 (Oct. 2013), 255–266. DOI:https://doi.org/10.11646/zootaxa.3731.2.6.