Génier, F. and Howden, H.F. 2014.

Onthophagus fragosus n.sp. a second endemic species of Onthophagus Latreille from Cuba (Coleoptera : Scarabaeidae, Scarabaeinae)

. Zootaxa. 3795, 5 (May 2014), 597–599. DOI:https://doi.org/10.11646/zootaxa.3795.5.10.