López-García, M.M., Gasca-Álvarez, H.J. and Amat-García, G. 2014. A new species of Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini), with a key to the species in Colombia. Zootaxa. 3869, 5 (Oct. 2014), 579–584. DOI:https://doi.org/10.11646/zootaxa.3869.5.8.