Nakahara, S., Fratello, S.A. and Harvey, D.J. 2014. A new species of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae: Satyrini) from Mount Roraima, Guyana. Zootaxa. 3881, 3 (Nov. 2014), 291–300. DOI:https://doi.org/10.11646/zootaxa.3881.3.7.