Bickel, D.J. 2015.

The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus

. Zootaxa. 4020, 1 (Sep. 2015), 169–182. DOI:https://doi.org/10.11646/zootaxa.4020.1.7.