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Lobo, J.M., Jiménez-Ruiz, Y., Chehlarov, E., Guéorguiev, B., Petrova, Y., Král, D., Alonso-Zarazaga, M. Ángel and Verdú, J.R. 2015. The classification and phylogenetic status of Jekelius (Reitterius) punctulatus (Jekel, 1866) and Jekelius (Jekelius) brullei (Jekel, 1866) (Coleoptera: Geotrupidae) using molecular data. Zootaxa. 4040, 2 (Nov. 2015), 187–203. DOI:https://doi.org/10.11646/zootaxa.4040.2.5.