Ahmed, Q., Thandar, A.S. and Ali, Q.M. 2020.

Holothuria (Lessonothuria) insignis Ludwig, 1875 (formally resurrected from synonymy of H. pardalis Selenka, 1867) and Holothuria (Lessonothuria) lineata Ludwig, 1875—new additions to the sea cucumber fauna of Pakistan, with a key to the subgenus Lessonothuria Deichmann (Echinodermata: Holothuroidea)

. Zootaxa. 4767, 2 (Apr. 2020), 307–318. DOI:https://doi.org/10.11646/zootaxa.4767.2.6.