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Benine, R.C. and Lopes, G.A.M. 2008. A new species of Hyphessobrycon (Characiformes: Characidae) from rio Portuguesa, rio Orinoco basin, Venezuela. Zootaxa. 1747, 1 (Apr. 2008), 61–68. DOI:https://doi.org/10.11646/zootaxa.1747.1.3.