Castroviejo-Fisher, S., Guayasamin, J.M. and Kok, P.J. 2009. Species status of Centrolene lema Duellman and Señaris, 2003 (Amphibia: Centrolenidae) revealed by Integrative Taxonomy. Zootaxa. 1980, 1 (Jan. 2009), 16–28. DOI:https://doi.org/10.11646/zootaxa.1980.1.2.