Bartels, P.J., Pilato, G., Lisi, O. and Nelson, D.R. 2009. Macrobiotus (Eutardigrada, Macrobiotidae) from the Great Smoky Mountains National Park, Tennessee/North Carolina, USA (North America): two new species and six new records. Zootaxa. 2022, 1 (Feb. 2009), 45–57. DOI:https://doi.org/10.11646/zootaxa.2022.1.4.