Lötters, S., Schmitz, A., Reichle, S., Rödder, D. and Quennet, V. 2009. Another case of cryptic diversity in poison frogs (Dendrobatidae: Ameerega)—description of a new species from Bolivia. Zootaxa. 2028, 1 (Mar. 2009), 20–30. DOI:https://doi.org/10.11646/zootaxa.2028.1.2.