Rödel, M.-O., Schmitz, A., Pauwels, O.S. and Böhme, W. 2004. Revision of the genus Werneria Poche, 1903, including the descriptions of two new species from Cameroon and Gabon (Amphibia: Anura: Bufonidae). Zootaxa. 720, 1 (Nov. 2004), 1–28. DOI:https://doi.org/10.11646/zootaxa.720.1.1.