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Perkins, P.D. 2004. A revision of the South African endemic humicolous beetle genus Nucleotops Perkins and Balfour-Browne (Coleoptera: Hydraenidae). Zootaxa. 559, 1 (Jul. 2004), 1–12. DOI:https://doi.org/10.11646/zootaxa.559.1.1.