Skelley, P.E. , Smith, A.B. and Mora-Aguilar, E.F. 2022.

A review of the flightless genus Nanotermitodius Howden, 2003 (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini)

. Zootaxa. 5200, 4 (Oct. 2022), 355–364. DOI:https://doi.org/10.11646/zootaxa.5200.4.4.