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Shapovalov, A.M. and Bagaturov, M.F. 2023. A review of the subgenus Mesolethrus Nikolajev, 2003 of the genus Lethrus Scopoli, 1777 (Coleoptera: Geotrupidae: Lethrinae) with description of a new species. Zootaxa. 5351, 4 (Sep. 2023), 435–452. DOI:https://doi.org/10.11646/zootaxa.5351.4.2.