[1]
Guayasamin, J.M., Bonaccorso, E., Duellman, W.E. and Coloma, L.A. 2010. Genetic differentiation in the nearly extinct harlequin frogs (Bufonidae: Atelopus), with emphasis on the Andean Atelopus ignescens and A. bomolochos species complexes. Zootaxa. 2574, 1 (Aug. 2010), 55–68. DOI:https://doi.org/10.11646/zootaxa.2574.1.2.