Johnson, C. B.; S. H., E. Q.; Anuar, S.; Muin, M. A.; Wood, Jr., P. L.; Grismer, J. L.; Greer, L. F.; Onn, C. K.; Ahmad, N.; Bauer, A. M.; Grismer, L. L. Phylogeography, Geographic Variation, and Taxonomy of the Bent-Toed Gecko Cyrtodactylus Quadrivirgatus Taylor, 1962 from Peninsular Malaysia With the Description of a New Swamp Dwelling Species. Zootaxa 2012, 3406, 39–58.