Pfingstl, T.; Schäffer, S.; Ebermann, E.; Krisper, G. Differentiation Between Two Epilittoral Species, Scutovertex Arenocolus Spec. Nov. And Scutovertex Pilosetosus Polderman (Acari: Oribatida) from Different European Coasts. Zootaxa 2009, 2153, 35–54.