Vigle, G. O.; Coloma, L. A.; Santos, J. C.; Hernandez-Nieto, S.; Ortega-Andrade, H. M.; Paluh, D. J.; Read, M.

A New Species of Leucostethus (Anura: Dendrobatidae) from the Cordillera Mache-Chindul in Northwestern Ecuador, With Comments on Similar Colostethus and Hyloxalus

. Zootaxa 2020, 4896, 342–372.