Jansen, M., & Schulze, A. (2012). Molecular, morphology and bioacoustic data suggest Bolivian distribution of a large species of the Leptodactylus pentadactylus group (Amphibia: Anura: Leptodactylidae). Zootaxa, 3307(1), 35–47. https://doi.org/10.11646/zootaxa.3307.1.2