Cardoso, Y. P., Almirón, A., Casciotta, J., Aichino, D., Lizarralde, M. S., & Montoya-Burgos, J. I. (2012). Origin of species diversity in the catfish genus Hypostomus (Siluriformes: Loricariidae) inhabiting the Paraná river basin, with the description of a new species. Zootaxa, 3453(1), 69–83. https://doi.org/10.11646/zootaxa.3453.1.5