Nunes, R. V., & Vaz-De-Mello, F. Z. (2016). A new species of Holocephalus Hope from Paraguay (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini s. str.). Zootaxa, 4136(2), 397–400. https://doi.org/10.11646/zootaxa.4136.2.12